Jeffrey A Fletcher

The evolution of altruism: game theory in multilevel selection and inclusive fitness.

Martin Zwick, J. Fletcher, M. Zwick, 2007, Journal of theoretical biology.

A simple and general explanation for the evolution of altruism

Michael Doebeli, J. Fletcher, M. Doebeli, 2009, Proceedings of the Royal Society B: Biological Sciences.

Hamilton's rule in multi-level selection models.

Burton Simon, Michael Doebeli, J. Fletcher, 2012, Journal of theoretical biology.

Strong altruism can evolve in randomly formed groups.

Martin Zwick, J. Fletcher, M. Zwick, 2004, Journal of theoretical biology.