Sandra K. Memenis

Rank-related partner choice in the fission–fusion society of the spotted hyena (Crocuta crocuta)

Kay E. Holekamp, K. Holekamp, Jennifer E. Smith, 2007, Behavioral Ecology and Sociobiology.

Evolutionary forces favoring intragroup coalitions among spotted hyenas and other animals

Kay E. Holekamp, K. Holekamp, Jennifer E. Smith, 2010 .