Ricardo Mallarino

Molecular systematics of the butterfly genus Ithomia (Lepidoptera: Ithomiinae): a composite phylogenetic hypothesis based on seven genes.

E. Bermingham, C. Jiggins, K. Willmott, 2005, Molecular phylogenetics and evolution.

The phylogenetic pattern of speciation and wing pattern change in neotropical Ithomia butterflies (Lepidoptera: nymphalidae).

E. Bermingham, C. Jiggins, K. Willmott, 2006 .

North Andean origin and diversification of the largest ithomiine butterfly genus

J. Mallet, M. Elias, G. Lamas, 2017, Scientific Reports.