A LONG‐SNOUTED DYROSAURID (CROCODYLIFORMES, MESOEUCROCODYLIA) FROM THE PALEOCENE OF MOROCCO: PHYLOGENETIC AND PALAEOBIOGEOGRAPHIC IMPLICATIONS

Abstract:  New material of a long‐snouted dyrosaurid has been discovered in the Paleocene of Morocco. It consists of a well‐preserved skull with embedded mandible and four dorsal vertebrae. The particularly elongate snout, proportionally the longest of all known dyrosaurids, allows precise identification of this material as Atlantosuchus coupatezi Buffetaut, 1979a , and presentation of an emended diagnosis for this species previously known only from a mandibular symphysis. A phylogenetic analysis of the dyrosaurids indicates a close relationship between A. coupatezi and Rhabdognathus. It also confirms a previous hypothesis that Congosaurus is distinct from Hyposaurus. It is more closely related to Atlantosuchus than Hyposaurus. The analysis also allows palaeobiogeographic interpretations to be made. Dyrosaurids ranged from North Africa to other areas. They were rare during the Maastrichtian and endemic to each continent at this time. Competition with large marine reptiles, such as mosasaurs, limited their dispersal during the Late Cretaceous. The disappearance of these rivals during the ‘K‐T crisis’ enabled their diversification and widespread dispersal during the Paleocene, with the same genera present on several continents.

[1]  S. Jouve TAXONOMIC REVISION OF THE DYROSAURID ASSEMBLAGE (CROCODYLIFORMES: MESOEUCROCODYLIA) FROM THE PALEOCENE OF THE IULLEMMEDEN BASIN, WEST AFRICA , 2007, Journal of Paleontology.

[2]  S. Jouve,et al.  A new species of Dyrosaurus (Crocodylomorpha, Dyrosauridae) from the early Eocene of Morocco: phylogenetic implications , 2006 .

[3]  S. Jouve,et al.  New material of Argochampsa krebsi (Crocodylia: Gavialoidea) from the Lower Paleocene of the Oulad Abdoun Basin (Morocco): phylogenetic implications , 2006 .

[4]  S. Jouve,et al.  A SHORT‐SNOUTED DYROSAURID (CROCODYLIFORMES, MESOEUCROCODYLIA) FROM THE PALAEOCENE OF MOROCCO , 2005 .

[5]  S. Jouve A new description of the skull of Dyrosaurus phosphaticus (Thomas, 1893) (Mesoeucrocodylia: Dyrosauridae) from the Lower Eocene of North Africa , 2005 .

[6]  S. Jouve,et al.  A PRIMITIVE MARINE GAVIALOID FROM THE PALEOCENE OF MOROCCO , 2004 .

[7]  M. O'Leary,et al.  A DYROSAURID CROCODYLIFORM BRAINCASE FROM MALI , 2002, Journal of Paleontology.

[8]  V. Gallo,et al.  Vertebrate assemblage from the Maria Farinha Formation after the K-T boundary , 2001 .

[9]  James M. Clark,et al.  Ancient marine reptiles , 1998 .

[10]  K. Bremer,et al.  BRANCH SUPPORT AND TREE STABILITY , 1994 .

[11]  A. D. Walker A revision of the Jurassic reptile Hallopus victor (Marsh), with remarks on the classification of crocodiles , 1970 .

[12]  G. Miller,et al.  Second Bibliography and Catalogue of the Fossil Vertebrata of North America , 1930 .

[13]  E. L. Troxell Hyposaurus, a marine crocodilian , 1925 .

[14]  Efic Jlntnan,et al.  Reversion , 1872, The Indian medical gazette.

[15]  S. Jouve,et al.  New material of Argochampsa krebsi (Eusuchia: Gavialoidea) and phylogenetic implications. , 2006 .

[16]  S. Jouve,et al.  A new dyrosaurid crocodyliform from the Palaeocene of Morocco and a phylogenetic analysis of Dyrosauridae , 2005 .

[17]  D. Schwarz,et al.  Congosaurus bequaerti, a Paleocene Dyrosaurid (Crocodyliformes; Mesoeucrocodylia) from Landana (Angola) , 2004 .

[18]  Stéphane Jouve Etude des crocodyliformes fini Crétacé-Paléogène du bassin des Oulad Abdoun (Maroc) et comparaison avec les faunes africaines contemporaines : systématique, phylogénie et paléobiogéographie , 2004 .

[19]  D. Swofford PAUP*: Phylogenetic analysis using parsimony (*and other methods), Version 4.0b10 , 2002 .

[20]  J. Dobie,et al.  The Marine Crocodilian Hyposaurus in North America , 1997 .

[21]  S. Hua A propos d'un crâne de Dyrosaurus phosphaticus (Thomas, 1893), (Crocodylia, Mesosuchia) : Contribution à l'interprétation paléoécologique et biogéographique des Dyrosauridae : Jurassic - Triassic vertebrates , 1995 .

[22]  B. Sigé,et al.  Découverte de vertébrés aquatiques présumés paléocènes dans les Andes septentrionales de Bolivie (Rio Suches, synclinorium de Putina) , 1987 .

[23]  É. Buffetaut Les Dyrosauridae (Crocodylia, Mesosuchia) des phosphates de l'Eocène inférieur de Tunisie : Dyrosaurus, Rhabdognatus, Phospliatosaurus , 1978 .

[24]  Archosaur phylogeny and the relationships of the Crocodyli a , 2022 .