Ingrowth, survival and height growth of small trees in uneven-aged Picea abies stands in southern Finland

The purpose of the study was to analyse height growth, mortality, and ingrowth of individual small-sized trees in uneven-aged spruce-dominated stands. It was based on experimental data from 16 stands for a 15-year observation period including four measurements with a 5-year interval. In the data of this study, the heights of small-sized trees varied from 0.1 to about 9 m. Results showed that the growth of small trees was rather slow, particularly in the smallest size classes. With average growth rates it would take about 60 years for a freshly emerged spruce germinant to achieve 1.3 m in height. The stand density, site quality and selection cuttings affected the growth of small-sized spruces. Average five-year mortality rates for spruce, birch and pine were 17.0%, 40.9% and 33.9%, respectively. Annual ingrowth rates with the threshold height of 1.3 m for the three species were on average 30.4, 2.8 and less than 0.1 trees per hectare, respectively. Even if, a selection cutting of modest intensity (25% of basal area removed on average) seemed to have accelerated the growth of small spruces, it is recommended that more intensive harvestings be applied to enhance the survival and growth of small spruces. It is also concluded that shade intolerant species like birch and pine do not seem to be capable of developing into viable undergrowth in spruce selection stands with their current levels of stand density.

[1]  Timo Pukkala,et al.  Growth and yield models for uneven-sized forest stands in Finland. , 2009 .

[2]  L. Lundqvist Changes in the stand structure on permanent Picea abies plots managed with single‐tree selection , 1993 .

[3]  Sauli Valkonen,et al.  Models for height development of Norway spruce and Scots pine advance growth after release in southern Finland. , 1993 .

[4]  Kristina Nilson Regeneration dynamics in uneven-aged Norway spruce forests with special emphasis on single-tree selection , 2001 .

[5]  K. Hanssen Effects of Seedbed Substrates on Regeneration of Picea abies from Seeds , 2002 .

[6]  O. Tahvonen OPTIMAL CHOICE BETWEEN EVEN‐ AND UNEVEN‐AGED FORESTRY , 2008 .

[7]  Timo Pukkala,et al.  Optimizing the structure and management of uneven-sized stands of Finland , 2010 .

[8]  K. Eerikäinen,et al.  Models for the regeneration establishment and the development of established seedlings in uneven-aged, Norway spruce dominated forest stands of southern Finland , 2007 .

[9]  Terje Gobakken,et al.  T: A forest simulator for bioeconomic analyses based on models for individual trees , 2008 .

[10]  Hubert Hasenauer,et al.  Sustainable forest management : growth models for Europe , 2006 .

[11]  Spatial distribution of injuries to Norway spruce advance growth after selection harvesting , 2001 .

[12]  R Core Team,et al.  R: A language and environment for statistical computing. , 2014 .

[13]  Timo Pukkala,et al.  A stochastic spatial regeneration model for Pinus sylvestris , 1992 .

[14]  T. Saksa,et al.  Dynamics of seedling establishment and survival in uneven-aged boreal forests , 2011 .

[15]  P. Wikström,et al.  A solution method for uneven-aged management applied to Norway spruce , 2000 .

[16]  E. Valinger,et al.  Stand development after different thinnings in two uneven-aged Picea abies forests in Sweden , 2007 .

[17]  D. Chrimes,et al.  Overstorey density influence on the height of Picea abies regeneration in northern Sweden , 2005 .

[18]  L. Lundqvist Some notes on the regeneration of Norway spruce on six permanent plots managed with single-tree selection , 1991 .

[19]  K. Hanssen,et al.  Performance of Sown and Naturally Regenerated Picea abies Seedlings Under Different Scarification and Harvesting Regimens , 2003 .

[20]  K. Hanssen Natural regeneration of Picea abies on small clear-cuts in SE Norway , 2003 .

[21]  N. E. Dekatoff THE NATURAL REGENERATION OF SPRUCE , 1932 .

[22]  N. Lexerød Recruitment models for different tree species in Norway , 2005 .

[23]  Taneli Kolström,et al.  Modelling the development of an uneven‐aged stand of Picea abies , 1993 .

[24]  L. Lundqvist Simulation of Sapling Population Dynamics in Uneven-aged Picea abies Forests , 1995 .

[25]  H. Goldstein Multilevel Statistical Models , 2006 .

[26]  Timo Pukkala,et al.  Optimal management of uneven-aged Norway spruce stands , 2010 .

[27]  D. Fjeld,et al.  Injuries After Selection Harvesting in Multi-Stored Spruce Stands– The Influence of Operating Systems and Harvest Intensity , 2013 .

[28]  C. Karlsson,et al.  Influence of Shelterwood Density on Survival and Height Increment of Picea abies Advance Growth , 2000 .

[29]  Hubert Hasenauer,et al.  Modeling Forest Regeneration , 2006 .

[30]  A. Cajander,et al.  Theory of forest types , 1926 .

[31]  T. Saksa Regeneration Process from Seed Crop to Saplings - a Case Study in Uneven- Aged Norway Spruce-Dominated Stands in Southern Finland , 2004 .

[32]  L. Lundqvist,et al.  Influence of local stand basal area on density and growth of regeneration in uneven‐aged Picea abies stands , 1996 .

[33]  D. Maguire,et al.  Relationship between seedbed properties and the emergence of spruce germinants in recently cut Norway spruce selection stands in Southern Finland , 2005 .

[34]  John McGregor,et al.  Ecosystems , 2009, J. Object Technol..

[35]  K. Andreassen,et al.  Economic consequences of three silvicultural methods in uneven‐aged mature coastal spruce forests of central Norway , 2002 .

[36]  V. Carey,et al.  Mixed-Effects Models in S and S-Plus , 2001 .

[37]  J. Heikkinen,et al.  Damage to saplings in mechanized selection cutting in uneven-aged Norway spruce stands , 2011 .

[38]  A. Cescatti,et al.  Early response of Pinus sylvestris and Picea abies seedlings to an experimental canopy gap in a boreal spruce forest , 2003 .

[39]  Sreenivasan Ravi,et al.  Multilevel Statistical Models, 3rd edn , 2005 .