Spider crabs (Decapoda: Brachyura: Majoidea) from the upper Eocene of south Pyrenees (Huesca, Spain)

Majoidea are one of the most plesiomorphic clades of Eubrachyura and display a huge diversity in modern ecosystems. We describe one new fossil genus and three new species including Eoparanaxia eocenica n. gen. n. sp., Planobranchia elongata n. sp., Spinirostrimaia echinata n. sp.; and one indeterminate species tentatively assigned to Macrocheira sp. from the Pamplona Marls Formation (upper Eocene, southern Pyrenees, Spain). All of them are new or first reported from the Iberian Peninsula. This shows a highly diversified fauna associated with prodelta clays that favoured preservation of decapods and other invertebrates. Cluster analyses based on Jaccard and Raup-Crick coefficients of Eocene Majidae suggest close affinities of Iberia with other European basins. Fernando A. Ferratges. Departamento de Ciencias de la Tierra-IUCA, Universidad de Zaragoza, Zaragoza E-50009, Spain. fer.afk87@gmail.es orcid.org/0000-0002-9532-6972 Josep Lluis Domínguez. Padre Manjón, 12. 50010 Zaragoza, Spain. jl.domin@hotmail.com Àlex Ossó. Llorenç de Villalonga, 17B, 1-1 43007 Tarragona, Catalonia. aosso@tinet.cat orcid.org/0000-0003-2528-9915 Samuel Zamora. Instituto Geológico y Minero de España (IGME-CSIC), Residencia CSIC, Campus Aula Dei, Av. Montañana 1005, 50059 Zaragoza, Spain. s.zamora@igme.es orcid.org/0000-0002-3917-4628

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