Two New Species in the Tillandsia utriculata Complex (Bromeliaceae)

Two new species in the Tillandsia utriculata (L.) L. complex (Bromeliaceae) from Mexico are proposed. Both are herein described and illus- trated with their affinities discussed. The first new entity, T. aesii I. Ramo´rez & Carnevali from the states of Jalisco, Guerrero, and Oaxaca, is similar to T. makoyana Baker, but the new taxon has an open rosette with leaves that are green adaxially and white lepidote abaxially (vs. a funnelform rosette with gray leaves on both surfaces in T. makoyana), actinomor- phic flowers with a light apple-green corolla (vs. zygomorphic with a purple to light purple corolla). The second taxon proposed here, T. pinicola I. Ramo´rez & Carnevali, from the state of Oaxaca, is also similar to T. makoyana, but is a smaller plant with a more compact growth habit and leaves with proportionally shorter leaf blades that abruptly attenuate from a broad sheath (vs. gradually attenuate into the proportionally longer blade) into a sub-acicular apex. It is also characterized by typically growing on pines, hence the epithet. RESUMEN . Dos nuevas especies en el complejo Tillandsia utriculata (L.) L. (Bromeliaceae) de Mexico se proponen como nuevas. Ambas se describen, se discuten sus afinidades y se documentan con ilustraciones y fotografo´as. La primera especie T. aesii I. Ramo´rez & Carnevali de los estados de Jalisco, Guerrerro y Oaxaca, es similar a T. makoyana Baker pero la nueva especie tiene rosetas abiertas con hojas verdes adaxialmente y blanco lepidotas abaxialmente (vs. rosetas en forma de embudo con hojas grises en ambas superficies en T. makoyana), flores actinomor- ficas con corola verde manzana claro (vs. flores zigomorfas y con petalos purpura o purpura claro). La segunda especie, T. pinicola I. Ramo´rez & Carnevali, del estado de Oaxaca, es tambien similar a T. makoyana pero es una planta mas pequena con un habito de crecimiento mas compacto y hojas propor- cionalmente con laminas foliares mas cortas las cuales se atenuan abruptamente en una vaina ancha (vs. vainas atenuandose gradualmente en laminas foliares proporcionalmente mas largas) con un apice acicular. Esta nueva especie tambien se caracteriza por crecer en pinos, de allo´ su epo´teto. The genus Tillandsia L. was classified in seven subgenera by Mez (1896) and later supported by Smith and Downs (1977) in their Flora Neotropica treatment of the entire Bromeliaceae. The high levels of homoplasy in several morphological characters (such as petal appendages) considered important in traditional classifications (including that of Smith and Downs) have blurred our understanding of the limits of the supraspecific groups within the genus. Some of the