First Precise Locality Data for Onthophagus atriglabrus Howden and Gill and New State Record for Onthophagus anewtoni Howden and Génier (Coleoptera: Scarabaeidae: Scarabaeinae) in Mexico

The subfamily Scarabaeinae is one of the best studied and most diversified groups of Coleoptera, with ca. 6,200 species, 267 genera (Tarasov and Génier 2015), and 11 tribes (sensu Bouchard et al. 2011). This diversity is mainly concentrated in the tropics, at elevations ranging from the sea level to 3,500 m (Morón 2003). In Mexico, there are 274 known scarabaeine species (Morón 2003; Howden and Génier 2004; Kohlmann and Solı́s 2006a, b; Delgado and Kohlmann 2007; Vaz-de-Mello 2008; Génier 2009; Halffter and Halffter 2009; Edmonds and Zidek 2012; Gómez 2013; González-Alvarado and Vaz-de-Mello 2014; Halffter et al. 2015; Mora-Aguilar and Delgado 2015; Arriaga-Jiménez et al. 2016; Moctezuma et al. 2016, 2017, 2018; Moctezuma and Halffter 2017; Kohlmann et al. 2018 a, b; Kohlmann and Vaz-de-Mello 2018). Among Mexican states, Chiapas, with 119 species, has the second highest species richness (SánchezHernández et al. 2018). However, the greatest