Modes of Selection

At the lowest level is selection among genes in mere capacity to persist and in mechanism of exact duplication. It seems probable, as suggested by Darwin (cf. Hardin 1950) and developed by Haldane (1933) and Oparin (1938), that life originated in an organic soup that could only have accumulated in the prior absence of life and that was one that provided all necessary metabolites. It is hardly probable that the chemical basis for gene persistence and duplication appeared full fledged. We may conclude with Blum (1951) that evolution took the form at first of decreasing mutability. Along with this may have occurred the evolution of stable genic association in cells, and mechanisms of exact duplication of the entire system by mitosis. *Address of the President, Society for the Study of Evolution, delivered at the

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