OpposingBMPandEGF signallingpathways convergeon theTGF-b familymediatorSmad 1

Acknowledgements. We thank J. Nüsslein-Volhard, Y. Chen and G. Struhl for fly stocks; R. Burke, C. Dahmann, N. Méthot and particularly G. Struhl for comments on the manuscript; F. Cifuentes, D. Nellen and M. Zecca for help with the clonal analysis; and J. Modolell for support. I.R. was the recipient of an EMBO fellowship. This work was supported by the Swiss National Science Foundation, the Direccion general de Investigacion Cientifica y Tecnica, and the Fundacion Ramon Areces.

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