Concepts of Scale in Landscape Ecology

This chapter seeks to evaluate the degree to which issues of spatial scale and scaling problems have been incorporated into different research problems in landscape ecology and to identify research methods applicable to problems of spatial scale. Then, the study identifies research foci in which scale has been relatively little explored, in part to determine if some questions have been largely divorced from questions of scale or whether appropriate conceptual or methodological approaches are lacking. The results may shed some light on to what degree, and in what subdisciplines within landscape ecology, the “paradigm shift” noted by Golley (1989) has been embraced. Finally, a standard vocabulary is proposed for issues of scale in the field of Landscape Ecology.

[1]  Thomas M. Smith,et al.  Modeling the Potential Response of Vegetation to Global Climate Change , 1992 .

[2]  J. R. Wallis,et al.  Some ecological consequences of a computer model of forest growth , 1972 .

[3]  R. Whittaker Evolution and measurement of species diversity , 1972 .

[4]  Monica G. Turner,et al.  Methods to evaluate the performance of spatial simulation models , 1989 .

[5]  G. Whitney Vegetation-Site Relationships in the Presettlement Forests of Northeastern Ohio , 1982, Botanical Gazette.

[6]  Arild O. Gautestad,et al.  Are home ranges fractals? , 2004, Landscape Ecology.

[7]  David J. Maguire,et al.  Geographical Information Systems , 1993 .

[8]  P. White,et al.  Scale Dependence and the Species-Area Relationship , 1994, The American Naturalist.

[9]  C. Goodess,et al.  The Development of Regional Climate Scenarios and the Ecological Impact of Greenhouse Gas Warming , 1992 .

[10]  L. Penev Qualitative and quantitative spatial variation in soil wire-worm assemblages in relation to climatic and habitat factors , 1992 .

[11]  L. Bersier,et al.  Bird assemblages in mosaic forests: the relative importance of vegetation structure and floristic composition along the successional gradient , 1994 .

[12]  Robert E. Ulanowicz,et al.  Scale and Biodiversity Policy: A Hierarchical Approach , 1994 .

[13]  Cliff R. Hupp,et al.  Bottomland Vegetation Distribution along Passage Creek, Virginia, in Relation to Fluvial Landforms , 1985 .

[14]  Raupach,et al.  Challenges in Linking Atmospheric CO2 Concentrations to Fluxes at Local and Regional Scales , 1992 .

[15]  Kathryn E. Freemark,et al.  Assessing effects of agriculture on terrestrial wildlife: developing a hierarchical approach for the US EPA , 1995 .

[16]  W. Marcus Copper dispersion in ephemeral stream sediments , 1987 .

[17]  K. Matthews,et al.  Climate change impact on distribution and abundance of wildlife species: an analytical approach using GIS. , 1994, Environmental pollution.

[18]  R. Macarthur,et al.  The Theory of Island Biogeography , 1969 .

[19]  Earl D. McCoy,et al.  The Statistics and Biology of the Species-Area Relationship , 1979, The American Naturalist.

[20]  L. Price Vegetation, Microtopography, and Depth of Active Layer on Different Exposures in Subarctic Alpine Tundra. , 1971, Ecology.

[21]  Mike Rees,et al.  5. Statistics for Spatial Data , 1993 .

[22]  D. Currie,et al.  Species-energy theory and patterns of species richness: I. Patterns of bird, angiosperm, and mammal species richness on islands , 1993 .

[23]  A. Shmida,et al.  Plant species diversity along a Mediterranean-desert gradient and its correlation with interannual rainfall fluctuations , 1992 .

[24]  J. Koricheva,et al.  The relationship between abundance and performance of Eriocrania miners in the field: effects of the scale and larval traits studied , 1994 .

[25]  Antoine Morin,et al.  Interpreting ecological patterns generated through simple stochastic processes , 1991, Landscape Ecology.

[26]  Steven K. Friedman,et al.  Assessing landscape dynamics in a protected area , 1992 .

[27]  J. Travis,et al.  Does the Spatial Scale of Experimentation Matter? A Test with Tadpoles and Dragonflies , 1992 .

[28]  Douglas Stow,et al.  Assessing the relationship between spectral vegetation indices and shrub cover in the Jornada Basin, New Mexico , 1993 .

[29]  T. Mitchell-Olds,et al.  Does environmental variation maintain genetic variation? A question of scale. , 1992, Trends in ecology & evolution.

[30]  K. M. O'Neill Temporal and spatial dynamics of predation in a robber fly (Efferia staminea) population (Diptera: Asilidae) , 1992 .

[31]  W. Hargrove,et al.  Landscape-scale heterogeneity in lodgepole pine serotiny , 1994 .

[32]  W. Calder,et al.  ECOLOGICAL SCALING: MAMMALS AND BIRDS , 1983 .

[33]  David M. Stoms Scale Dependence of Species Richness Maps , 1994 .

[34]  Robert B. Jackson,et al.  Geostatistical Patterns of Soil Heterogeneity Around Individual Perennial Plants , 1993 .

[35]  ECOGIS/ECOSAD: A methodology for the biophysical environmental assessment within the planning process , 1993 .

[36]  G Sugihara,et al.  Applications of fractals in ecology. , 1990, Trends in ecology & evolution.

[37]  A. Magurran,et al.  Biological diversity : the coexistence of species on changing landscapes , 1994 .

[38]  M. Goodchild,et al.  The fractal properties of topography: A comparison of methods , 1992 .

[39]  Shelley A. Hinsley,et al.  Habitat and landscape factors influencing the presence of individual breeding bird species in woodland fragments , 1995 .

[40]  M. Claussen,et al.  LANDSCAPE VARIABILITY AND SURFACE FLUX PARAMETERIZATION IN CLIMATE MODELS , 1995 .

[41]  S. Running Estimating Terrestrial Primary Productivity by Combining Remote Sensing and Ecosystem Simulation , 1990 .

[42]  W. Kemp Rangeland Grasshopper (Orthoptera: Acrididae) Community Structure: A Working Hypothesis , 1992 .

[43]  N. Stenseth,et al.  Ecological mechanisms and landscape ecology , 1993 .

[44]  B. Mandelbrot Fractal Geometry of Nature , 1984 .

[45]  I. Moore,et al.  Fractals, fractal dimensions and landscapes — a review , 1993 .

[46]  A. Ssymank,et al.  Proposal on a criteria system for a National Red Data Book of Biotopes , 1995, Landscape Ecology.

[47]  D. H. Knight,et al.  Fire Frequency and Subalpine Forest Succession Along a Topographic Gradient in Wyoming , 1981 .

[48]  Lawrence E. Band,et al.  Scale: Landscape attributes and geographical information systems , 1995 .

[49]  David Harvey,et al.  Pattern, Process, and the Scale Problem in Geographical Research , 1968 .

[50]  H. Gleason,et al.  On the Relation Between Species and Area , 1922 .

[51]  S. M. Glenn,et al.  Disturbances in tallgrass prairie: local and regional effects on community heterogeneity , 1992, Landscape Ecology.

[52]  J. M. Thomas,et al.  Multiple landscape scales: An intersite comparison , 2004, Landscape Ecology.

[53]  Frederick E. Petry,et al.  Principles and Applications , 1997 .

[54]  Hal Caswell,et al.  Community Structure: A Neutral Model Analysis , 1976 .

[55]  Neo D. Martinez,et al.  Scale and food-web structure: from local to global , 1995 .

[56]  Michael S. Gaines,et al.  Habitat Fragmentation and Movements of Three Small Mammals (Sigmodon, Microtus, and Peromyscus) , 1995 .

[57]  James S. Clark,et al.  Integration of ecological levels: individual plant growth, population mortality and ecosystem processes. , 1990 .

[58]  Fei Chen,et al.  Development and analysis of prognostic equations for mesoscale kinetic energy and mesoscale (subgrid scale) fluxes for large-scale atmospheric models , 1993 .

[59]  A. Ives,et al.  RESPONSE OF A PREDATOR TO VARIATION IN PREY DENSITY AT THREE HIERARCHICAL SCALES: LADY BEETLES FEEDING ON APHIDS' , 1993 .

[60]  Nathaniel A. Lifton,et al.  Tectonic, climatic and lithologic influences on landscape fractal dimension and hypsometry : implications for landscape evolution in the San Gabriel Mountains, California , 1992 .

[61]  B. Zimmerman,et al.  Relevance of the equilibrium theory of island biogeography and species-area relations to conservation with a case from Amazonia , 1986 .

[62]  Robert G. Bailey,et al.  The factor of scale in ecosystem mapping , 1985 .

[63]  J. Michaelsen,et al.  Regression Tree Analysis of satellite and terrain data to guide vegetation sampling and surveys , 1994 .

[64]  F. Giorgi,et al.  Use of output from high-resolution atmospheric models in landscape-scale hydrologic models: An assessment , 1993 .

[65]  John E. Estes,et al.  A remote sensing research agenda for mapping and monitoring biodiversity , 1993 .

[66]  N. Glasser,et al.  Evolution of large roches moutonnees. , 1992 .

[67]  B. Biswell,et al.  Northern Spotted Owls: Influence of Prey Base and Landscape Character , 1992 .

[68]  A. Cherrill A comparison of three landscape classifications and investigation of the potential for using remotely sensed land cover data for landscape classification , 1994 .

[69]  R. Forman,et al.  Patches and Structural Components for A Landscape Ecology , 1981 .

[70]  Kirk H. Stone Scale, Scale, Scale? , 1968 .

[71]  Michael N. DeMers Roadside ditches as corridors for range expansion of the western harvester ant (Pogonomyrmex occidentalis Cresson) , 1993, Landscape Ecology.

[72]  D. Kohn,et al.  Plant species richness: the effect of island size and habitat diversity , 1994 .

[73]  On the use of grain size in ecology , 1990 .

[74]  R. Sanderson,et al.  Predicting the distributions of plant species at the regional scale: a hierarchical matrix model , 1995, Landscape Ecology.

[75]  W. Baker Spatially heterogeneous multi-scale response of landscapes to fire suppression , 1993 .

[76]  M. Soulé,et al.  Conservation Biology: The Science of Scarcity and Diversity , 1986 .

[77]  Ann Henderson-Sellers,et al.  A Climate Modelling Primer , 1987 .

[78]  Steven W. Running,et al.  The effects of aggregating sub-grid land surface variation on large-scale estimates of net primary production , 1995, Landscape Ecology.

[79]  M. Palmer,et al.  The Coexistence of Species in Fractal Landscapes , 1992, The American Naturalist.

[80]  W. C. Clark,et al.  Scales of climate impacts , 1985 .

[81]  D. Strong,et al.  Does Spatial Scale Affect the Incidence of Density Dependence? A Field Test with Insect Parasitoids , 1991 .

[82]  Jared M. Diamond,et al.  THE ISLAND DILEMMA: LESSONS OF MODERN BIOGEOGRAPHIC STUDIES FOR THE DESIGN OF NATURAL RESERVES , 1975 .

[83]  R. Pielke,et al.  Nonlinear Influence of Mesoscale Land Use on Weather and Climate , 1991 .

[84]  D. Mccollin Avian distribution patterns in a fragmented wooded landscape (North Humberside, U.K.): the role of between-patch and within-patch structure , 1993 .

[85]  J. Woinarski,et al.  Patterns of species composition and reserve design for a fragmented estate: monsoon rainforests in the Northern Territory, Australia , 1995 .

[86]  Simon A. Levin,et al.  A Spatial Patch Dynamic Modeling Approach to Pattern and Process in an Annual Grassland , 1994 .

[87]  O. Vetaas Micro‐site effects of trees and shrubs in dry savannas , 1992 .

[88]  E. Grimm Fire and Other Factors Controlling the Big Woods Vegetation of Minnesota in the Mid‐Nineteenth Century , 1984 .

[89]  H. Delcourt,et al.  Presettlement landscape heterogeneity: Evaluating grain of resolution using General Land Office Survey data , 1996, Landscape Ecology.

[90]  J. Blondel,et al.  Do harlequin Mediterranean environments function as source sink for Blue Tits (Parus caeruleus L.)? , 1992, Landscape Ecology.

[91]  W. W. Pettapiece,et al.  Application of agro-ecological concepts and hierarchy theory in the design of databases for spatial and temporal characterisation of land and soil , 1993 .

[92]  C. Field,et al.  Scaling physiological processes: leaf to globe. , 1995 .

[93]  B. Danielson,et al.  Spatially Explicit Population Models: Current Forms and Future Uses , 1995 .

[94]  Peter B. Reich,et al.  Spatial Patterns and Succession in a Minnesota Southern‐Boreal Forest , 1995 .

[95]  C. W. Ramm,et al.  Landform-mediated differences in successional pathways among upland forest ecosystems in northwestern Lower Michigan. , 1987 .

[96]  K. Clarke Computation of the fractal dimension of topographic surfaces using the triangular prism surface area method , 1985 .

[97]  Y. Bergeron The Influence of Island and Mainland Lakeshore Landscapes on Boreal Forest Fire Regimes , 1991 .

[98]  R. Leemans,et al.  Pattern and process and the dynamics of forest structure: a simulation approach. , 1990 .

[99]  Robert V. O'Neill,et al.  Relating Patterns of Land‐Use Change to Faunal Biodiversity in the Central Amazon , 1994 .

[100]  P. Vitousek 10 – Global Dynamics and Ecosystem Processes: Scaling Up or Scaling Down? , 1993 .

[101]  Robert V. O'Neill,et al.  Spatio-Temporal Dispersal Strategies and Annual Plant Species Coexistence in a Structured Landscape , 1994 .

[102]  A. Bennett Microhabitat use by the long-nosed potoroo, Potorous tridactylus, and other small mammals in remnant forest vegetation, south-western Victoria , 1993 .

[103]  A. Bouchard,et al.  Riverine wetland vegetation: importance of small‐scale and large‐scale environmental variation , 1993 .

[104]  Elgene O. Box,et al.  Scale Effects in Landscape Studies , 1987 .

[105]  Pierre Dutilleul,et al.  Spatial Heterogeneity against Heteroscedasticity: An Ecological Paradigm versus a Statistical Concept , 1993 .

[106]  Anthony W King,et al.  Aggregating Fine-Scale Ecological Knowledge to Model Coarser-Scale Attributes of Ecosystems. , 1992, Ecological applications : a publication of the Ecological Society of America.

[107]  Yves Bergeron,et al.  Fractal dimension estimates of a fragmented landscape: sources of variability , 1994, Landscape Ecology.

[108]  John A. Wiens,et al.  Animal movement in heterogeneous landscapes : an experiment with Eleodes beetles in shortgrass prairie , 1992 .

[109]  C. L. Knight,et al.  Expansion of gallery forest on Konza Prairie Research Natural Area, Kansas, USA , 1994, Landscape Ecology.

[110]  Stephen T. Buckland,et al.  Statistical modelling of regional GIS data: an overview , 1993 .

[111]  G. Likens,et al.  Pattern and process in a forested ecosystem. , 1979 .

[112]  Randy Elliot Bennett,et al.  METHODS FOR EVALUATING THE PERFORMANCE OF SCHOOL PSYCHOLOGISTS , 1979 .

[113]  Neo D. Martinez Scale-Dependent Constraints on Food-Web Structure , 1994, The American Naturalist.

[114]  David J. Maguire,et al.  Geographical information systems : principles and applications , 1991 .

[115]  C. Loehle,et al.  Landscape habitat diversity: a multiscale information theory approach , 1994 .

[116]  M. Goodchild,et al.  The Fractal Nature of Geographic Phenomena , 1987 .

[117]  J. Sanderson,et al.  Spatial scale of aggregation in three acarine predator species with different degrees of polyphagy , 1993, Oecologia.

[118]  W. Post Uncertainties in the Terrestrial Carbon Cycle , 1993 .

[119]  C. Sandoval THE EFFECTS OF THE RELATIVE GEOGRAPHIC SCALES OF GENE FLOW AND SELECTION ON MORPH FREQUENCIES IN THE WALKING‐STICK TIMEMA CRISTINAE , 1994, Evolution; international journal of organic evolution.

[120]  M. Turner,et al.  WINTER HABITAT USE BY LARGE UNGULATES FOLLOWING FIRE IN NORTHERN YELLOWSTONE NATIONAL PARK , 1995 .

[121]  W. Ehmann Organization of Spider Assemblages on Shrubs: An Assessment of the Role of Dispersal Mode in Colonization , 1994 .

[122]  W. H. Blackburn,et al.  Partitioning small scale spatial variability of runoff and erosion on sagebrush rangeland , 1994 .

[123]  D. Montgomery,et al.  Channel Initiation and the Problem of Landscape Scale , 1992, Science.

[124]  Steven W. Running,et al.  Testing scale dependent assumptions in regional ecosystem simulations , 1994 .

[125]  I. Zonneveld,et al.  The land unit — A fundamental concept in landscape ecology, and its applications , 1989, Landscape Ecology.

[126]  J. Phillips Biogeomorphology and landscape evolution: The problem of scale , 1995 .

[127]  Robin L. Graham,et al.  Regional forest cover estimation via remote sensing: the calibration center concept , 1994, Landscape Ecology.

[128]  Paul G. Risser,et al.  Scales and global change. Spatial and temporal variability in biospheric and geospheric processes. , 1988 .

[129]  Jean E. McKendry,et al.  The role of geography in extending biodiversity gap analaysis , 1993 .

[130]  S S Stevens,et al.  On the Theory of Scales of Measurement. , 1946, Science.

[131]  Scott L. Collins,et al.  The hierarchical continuum concept , 1993 .

[132]  J. Briggs,et al.  Management Practices in Tallgrass Prairie: Large- and Small-Scale Experimental Effects on Species Composition , 1993 .

[133]  Francisco J. Artigas,et al.  Forty-eight years of landscape change on two contiguous Ohio landscapes , 1994, Landscape Ecology.

[134]  George L. Ball Ecosystem modeling with GIS , 1994 .

[135]  S. P. Mclaughlin,et al.  Floristic plant geography: the classification of floristic areas and floristic elements , 1994 .

[136]  Brendan Mackey,et al.  Predicting the potential distribution of rain‐forest structural characteristics , 1994 .

[137]  S. Lavorel,et al.  Small scale disturbances and the maintenance of species diversity in Mediterranean old fields , 1994 .

[138]  A. O. Nicholls,et al.  Selecting networks of reserves to maximise biological diversity , 1988 .

[139]  S. Levin The problem of pattern and scale in ecology , 1992 .

[140]  S. Stuart,et al.  A methodology for analyzing rare species distribution patterns utilizing GIS technology: The rare birds of Tanzania , 1989, Landscape Ecology.

[141]  D. Montgomery,et al.  Limits to Relief , 1995, Science.

[142]  A. Magurran Ecological Diversity and Its Measurement , 1988, Springer Netherlands.

[143]  J. Wiens Spatial Scaling in Ecology , 1989 .

[144]  G. Barrett,et al.  Landscape Ecology: Designing Sustainable Agricultural Landscapes , 1992 .

[145]  B. Kennedy A question of scale? , 1977 .

[146]  I. Rodríguez‐Iturbe,et al.  The relationship between catchment and hillslope properties: implications of a catchment evolution model , 1992 .

[147]  A. Taber,et al.  Ranging behaviour and activity patterns of two sympatric peccaries, Catagonus wagneri and Tayassu tajacu, in the Paraguayan Chaco , 1994 .

[148]  R. G. Wright,et al.  An Ecological Evaluation of Proposed New Conservation Areas in Idaho: Evaluating Proposed Idaho National Parks , 1994 .

[149]  Bruce T. Milne,et al.  Lessons from applying fractal models to landscape patterns , 1991 .

[150]  Robert V. O'Neill,et al.  Pattern, process, and predictability: the use of neutral models for landscape analysis , 1991 .

[151]  Robert H. Whittaker,et al.  Evolution of Species Diversity in Land Communities , 1977 .

[152]  Agnès Bégué,et al.  Radiative transfer in shrub savanna sites in Niger: preliminary results from HAPEX-Sahel. 2. Photosynthetically active radiation interception of the woody layer , 1994 .

[153]  L. Queen,et al.  Crane habitat evaluation using GIS and remote sensing , 1993 .

[154]  F. D. Pineda,et al.  Influence of landscape complexity and land management on woody plant diversity in northern Spain , 1994 .

[155]  J. Fox The problem of scale in community resource management , 1992 .

[156]  Monica G. Turner,et al.  Quantitative methods in landscape ecology : the analysis and interpretation of landscape heterogeneity , 1991 .

[157]  P. Clergeau Importance of multiple scale analysis for understanding distribution and for management of an agricultural bird pest , 1995 .

[158]  Harold A. Mooney,et al.  Remote Sensing of Biosphere Functioning , 1990 .

[159]  Mark A. White,et al.  Comparing Spatial Pattern in Unaltered Old-Growth and Disturbed Forest Landscapes. , 1993, Ecological applications : a publication of the Ecological Society of America.

[160]  W. Hargrove,et al.  Effects of fire on landscape heterogeneity in Yellowstone National Park, Wyoming , 1994 .

[161]  G. Barrett,et al.  Optimizing habitat fragmentation: an agrolandscape perspective , 1994 .

[162]  J. M. Thomas,et al.  Satellite imagery as a tool to evaluate ecological scale , 1992, Landscape Ecology.

[163]  R. B. Jackson,et al.  The Scale of Nutrient Heterogeneity Around Individual Plants and Its Quantification with Geostatistics , 1993 .

[164]  B. Milne Measuring the fractal geometry of landscapes , 1988 .

[165]  Landscape-ecological mapping of the Netherlands , 1991, Landscape Ecology.

[166]  J. Ellis,et al.  Landscape and climatic control of woody vegetation in a dry tropical ecosystem: Turkana District, Kenya , 1993 .

[167]  R. Webster,et al.  Kriging: a method of interpolation for geographical information systems , 1990, Int. J. Geogr. Inf. Sci..

[168]  P. Schippers,et al.  Foraging in a landscape mosaic: selection for energy and minerals in free-ranging cattle , 1994, Oecologia.

[169]  Z. Naveh Red Books for threatened Mediterranean landscapes as an innovative tool for holistic landscape conservation. Introduction to the western Crete Red Book case study , 1993 .

[170]  R. Gardner,et al.  Quantitative methods in landscape ecology: an introduction , 1991 .

[171]  A. S. Belward,et al.  Scale considerations in vegetation monitoring using AVHRR data , 1992 .

[172]  S. DeGloria,et al.  Mapping atrazine leaching potential with integrated environmental databases and simulation models , 1995 .

[173]  James H. Brown,et al.  Spatial Scaling of Species Composition: Body Masses of North American Land Mammals , 1991, The American Naturalist.

[174]  T. Herben,et al.  Small-scale variability as a mechanism for large-scale stability in mountain grasslands , 1993 .

[175]  Luis A. Bojórquez-Tapia,et al.  Identifying Conservation Priorities in Mexico Through Geographic Information Systems and Modeling , 1995 .

[176]  John A. Wiens,et al.  What is landscape ecology, really? , 1992, Landscape Ecology.

[177]  P. Greig-Smith,et al.  The Use of Random and Contiguous Quadrats in the Study of the Structure of Plant Communities , 1952 .

[178]  J. Reynolds,et al.  A Simulation Experiment to Quantify Spatial Heterogeneity in Categorical Maps , 1994 .

[179]  George R. Parker,et al.  Using an index of habitat patch proximity for landscape design , 1994 .

[180]  P. Burrough Fractal dimensions of landscapes and other environmental data , 1981, Nature.

[181]  R. Peters The Ecological Implications of Body Size , 1983 .

[182]  N. C. Kenkel Modeling Markovian Dependence in Populations of Aralia Nudicaulis , 1993 .

[183]  W. N. Beckon The effect of insularity on the diversity of land birds in the Fiji islands: implications for refuge design , 1993, Oecologia.

[184]  P. Colinvaux Amazon diversity in light of the paleoecological record , 1987 .

[185]  L. Heidt,et al.  Carbon trace gas fluxes along a successional gradient in the Hudson Bay lowland , 1994 .

[186]  R. Gardner,et al.  Quantitative Methods in Landscape Ecology , 1991 .

[187]  H. Gleason Species and Area , 1925 .

[188]  Frank B. Golley,et al.  Paradigm shift , 1989, Landscape Ecology.

[189]  C. S. Holling Cross-Scale Morphology, Geometry, and Dynamics of Ecosystems , 1992 .

[190]  W. Edmonds,et al.  Variance and Scale Influences on Classifying and Interpreting Soil Map Units1 , 1985 .

[191]  M. Hutchinson,et al.  Estimating fractal dimension of profiles: A comparison of methods , 1994 .

[192]  M. Crawley,et al.  Spatial density dependence in populations of a cynipid gall-former Andricus quercuscalicis , 1992 .

[193]  C. M. Cowell,et al.  Presettlement Piedmont Forests: Patterns of Composition and Disturbance in Central Georgia , 1995 .

[194]  D. Nir Regional geography considered from the systems' approach , 1987 .

[195]  Michael J. Conroy,et al.  Parameter Estimation, Reliability, and Model Improvement for Spatially Explicit Models of Animal Populations , 1995 .

[196]  J. Lundquist Disturbance profile—a measure of small-scale disturbance patterns in ponderosa pine stands , 1995 .

[197]  Valerie I. Cullinan,et al.  A comparison of quantitative methods for examining landscape pattern and scale , 1992, Landscape Ecology.

[198]  Lisa J. Graumlich,et al.  Holocene Variation in Spatial Scales of Vegetation Pattern in the Upper Great Lakes , 1993 .

[199]  Frank W. Davis,et al.  An information systems approach to the preservation of biological diversity , 1990, Int. J. Geogr. Inf. Sci..

[200]  M. Turner,et al.  Usefulness of spatially explicit population models in land management , 1995 .

[201]  Steven A. Wolf,et al.  Recasting alternative agriculture as a management model: The value of adept scaling , 1995 .

[202]  River/land ecotones: scales and patterns , 1995 .

[203]  A. Cooper Multivariate land class and land cover correlations in Northern Ireland , 1995 .

[204]  J. Bendix Scale, Direction, and Pattern in Riparian Vegetation-Environment Relationships , 1994 .

[205]  M. Rosenzweig Species Diversity Gradients: We Know More and Less Than We Thought , 1992 .

[206]  P. Lavelle,et al.  A Hierarchical Model for Decomposition in Terrestrial Ecosystems: Application to Soils of the Humid Tropics , 1993 .

[207]  Thomas B. Starr,et al.  Hierarchy: Perspectives for Ecological Complexity , 1982 .

[208]  Robert V. O'Neill,et al.  Patternandscale: statistics for landscape ecology , 1991 .

[209]  J. Schaefer Spatial patterns in taiga plant communities following fire , 1993 .

[210]  P. Rey Spatio‐Temporal Variation in Fruit and Frugivorous Bird Abundance in Olive Orchards , 1995 .

[211]  M. Heit,et al.  GIS applications in natural resources , 1996 .

[212]  R. Sharitz Comments of the editor , 2004, Wetlands Ecology and Management.

[213]  H. Shugart,et al.  Vegetation Dynamics and Global Change , 1993 .

[214]  S. Warwick,et al.  Genetic variation and structure at three spatial scales for Acer saccharum (sugar maple) in Canada and the implications for conservation , 1993 .

[215]  Richard G. Lathrop,et al.  Spatial variability of digital soil maps and its impact on regional ecosystem modeling , 1995 .

[216]  Emery R. Boose,et al.  Hurricane impacts to tropical and temperate forest landscapes , 1994 .

[217]  Monica G. Turner,et al.  Landscape Heterogeneity and Disturbance , 2011, Ecological Studies.

[218]  David G. Tarboton,et al.  A Physical Basis for Drainage Density , 1992 .

[219]  D. Boer Hierarchies and spatial scale in process geomorphology: a review , 1992 .

[220]  Bruce T. Milne,et al.  Spatial Aggregation and Neutral Models in Fractal Landscapes , 1992, The American Naturalist.

[221]  C. Johnston Material Fluxes Across Wetland Ecotones in Northern Landscapes. , 1993, Ecological Applications.

[222]  A. King Translating models across scales in the landscape , 1991 .

[223]  L. Aarssen Causes and consequences of variation in competitive ability in plant communities , 1992 .

[224]  Robert D. Holt,et al.  Linking Contemporary Vegetation Models with Spatially Explicit Animal Population Models , 1995 .

[225]  David C. Schneider,et al.  Quantitative Ecology: Spatial and Temporal Scaling , 1994 .

[226]  R. O'Neill A Hierarchical Concept of Ecosystems. , 1986 .

[227]  Harto Lindén,et al.  Forest fragmentation due to agriculture affects the reproductive success of the ground-nesting black grouse Tetrao tetrix , 1995 .

[228]  J. M. Cornelius,et al.  On Determining the Statistical Significance of Discontinuities with Ordered Ecological Data , 1991 .

[229]  D P Faith,et al.  Phylogenetic pattern and the quantification of organismal biodiversity. , 1994, Philosophical transactions of the Royal Society of London. Series B, Biological sciences.

[230]  A. Gentry,et al.  Changes in Plant Community Diversity and Floristic Composition on Environmental and Geographical Gradients , 1988 .

[231]  Mikhail A. Vedyushkin Fractal properties of forest spatial structure , 1994, Vegetatio.

[232]  S. Gould ALLOMETRY AND SIZE IN ONTOGENY AND PHYLOGENY , 1966, Biological reviews of the Cambridge Philosophical Society.