Four new Aphodius Illiger from pocket gopher burrows in Arizona, Utah, Kansas and Nebraska (Coleoptera: Scarabaeidae: Aphodiinae)

We describe four new species of winter-active Aphodius (sensu lato) from pocket gopher burrows in Arizona, Utah, Kansas, and Nebraska: A. caccabatus, A. paulseni, A. skillmani, and A. utopensis. Diagnostic characters of all four species are illustrated.