Regulation of neuroblast competence in Drosophila

that the reactive control layer is inactive. (6) Chaining through LTM sequences is achieved by biasing the LTM matching process. The competition among LTM segments takes place on a quantity m where m k l of segment l of sequence k is defined as: m k l ¼ c k l t k l ð5Þ where c k l is defined as dðE; E k l Þ using the definition of d provided in equation (4). t k l , t k l [ ½0; 1Š; is a dynamic threshold that provides a probabilistic mechanism for chaining through a LTM sequence. In case segment l of sequence k wins the competition by virtue of having the lowest value of m, it will reduce t of segment l þ 1, of k; t k lþ1 ; to a fixed value b,b [ [0; 1]. t k l relaxes to its default value of 1 according to t k l ðt þ 1Þ ¼ a T þ ð1 2 a T Þt k l ðtÞ; a T [ ½0; 1Š: The LTM segment that wins the competition will dominate the behavioural output of the overall system when its m is below a fixed threshold. In the present implementation, STM is a ring buffer with a fixed length of 25 segments, while the capacity of LTM is limited to 64 sequences. Emergence of simple-cell receptive field properties by learning a sparse code for natural images. coding in the visual cortex: a functional interpretation of some extra-classical receptive-field effects. brain substrates of aversive classical conditioning.

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