Revision of Late Tithonian charophytes from Lisbon and Sintra-Cascais (Portugal): taxonomy, biostratigraphy and palaeobiogeographical significance

The re-evaluation of the charophyte content of ‘Purbeckian’ sediments from several sections around Lisbon (Olelas and Brouco) and Sintra-Cascais (Murches, Atrozela and Malveira-Guincho) revealed new palaeofloral associations from the Late Tithonian of the South Lusitanian Basin (Portugal). These sections contain Globator rectispirale, G. aff. nurrensis, Nodosoclavator bradleyi, Clavator reidi, Clypeator cf. discordis, Porochara maxima, and newly described occurrences of P. jaccardi, Mesochara harrisi and nodosoclavatoroide utricles. These revised data reinforce the evidence for assigning most of the studied deposits to a Late Tithonian age, instead of the formerly accepted wider interval Tithonian to Early Berriasian (‘Purbeckian’). Population analysis and statistics were applied in order to better assess population variation of the different species. The results of this study are relevant as they contribute to improve the biostratigraphical definition of the ‘Purbeckian’ formations of Portugal and allow more accurate palaeobiogeographical interpretations within the central Tethyan domain, by comparison of the identified charophyte assemblages with documented Jurassic–Cretaceous transition charophytes from other regions.