Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi River drainages. Part 2: Lizards (Sauria), chelonians, and crocodiles

. —This work is the second report of the results obtained from a series of rapid biodiversity surveys of the upper Cuito, Cubango, Cuando, Zambezi, and Kwanza River basins in Angola, which were conducted between 2015 and 2019 in conjunction with the National Geographic Okavango Wilderness Project. The herpetofauna of this region are poorly documented and the results of these surveys help to address the knowledge gap regarding the conservation importance of this region. Here, an updated checklist is provided for the current and historical records of lizards, chelonians, and crocodiles from the southeastern region of Angola. A total of 369 new records were documented comprising 40 species, bringing the total number of recognized lizard, chelonian, and crocodile species in this region to 58. These surveys documented four new country records (i.e., Lygodactylus chobiensis , Agama armata , Pachydactylus wahlbergii , and Ichnotropis cf. grandiceps ) and increased the total number of reptile species known to occur in Angola (excluding snakes) from approximately 157 to 161. Finally, updated distribution maps for the whole country are provided for all of the species encountered in this study.

[1]  A. Schmitz,et al.  Rock island melody remastered: two new species in the Afroedura bogerti Loveridge, 1944 group from Angola and Namibia , 2022, Zoosystematics and Evolution.

[2]  W. Conradie,et al.  A new species of feather-tailed leaf-toed gecko, Kolekanos Heinicke, Daza, Greenbaum, Jackman, Bauer, 2014 (Squamata, Gekkonidae) from the poorly explored savannah of western Angola , 2022, ZooKeys.

[3]  A. Bauer,et al.  The Angolan bushveld lizards, genus Heliobolus Fitzinger, 1843 (Squamata: Lacertidae): Integrative taxonomy and the description of two new species , 2022, Vertebrate Zoology.

[4]  B. Jansen van Vuuren,et al.  Out of southern Africa: origins and cryptic speciation in Chamaeleo, the most widespread chameleon genus. , 2022, Molecular phylogenetics and evolution.

[5]  K. Tolley,et al.  An updated herpetofaunal species inventory of Iona National Park in southwestern Angola , 2022, Check List.

[6]  A. Bauer,et al.  All in all its just another branch in the tree: A new species of Acanthocercus Fitzinger, 1843 (Squamata: Agamidae), from Angola. , 2022, Zootaxa.

[7]  K. Tolley,et al.  Mistaken identity: challenges with specimen identification for morphologically conservative skinks (Trachylepis) leads to taxonomic error , 2022, African Journal of Herpetology.

[8]  A. Bauer,et al.  A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. , 2021, Zootaxa.

[9]  A. Bauer,et al.  How the African house gecko (Hemidactylus mabouia) conquered the world , 2021, Royal Society Open Science.

[10]  A. Schmitz,et al.  Rock island melody: A revision of the Afroedura bogerti Loveridge, 1944 group, with descriptions of four new endemic species from Angola , 2021 .

[11]  A. Bauer,et al.  A correction to a recent review of the genus Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from Angola, with the description of two additional species. , 2020, Zootaxa.

[12]  A. Bauer,et al.  A revision of the Dwarf Geckos, genus Lygodactylus (Squamata: Gekkonidae), from Angola, with the description of three new species. , 2020, Zootaxa.

[13]  A. Bauer,et al.  A review of the African snake-eyed skinks (Scincidae: Panaspis) from Angola, with the description of a new species. , 2020, Zootaxa.

[14]  A. Bauer,et al.  A review of the genus Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from Angola, with the description of two new species. , 2020, Zootaxa.

[15]  K. Tolley,et al.  Another Angolan Namib endemic species: a new Nucras Gray, 1838 (Squamata: Lacertidae) from south-western Angola , 2020 .

[16]  A. Bauer,et al.  A new species of Girdled Lizard (Squamata: Cordylidae) from the Serra da Neve Inselberg, Namibe Province, southwestern Angola. , 2019, Zootaxa.

[17]  A. Bauer,et al.  A species-level phylogeny of Trachylepis (Scincidae: Mabuyinae) provides insight into their reproductive mode evolution. , 2019, Molecular phylogenetics and evolution.

[18]  A. Bauer,et al.  Description of a new long-tailed skink (Scincidae: Trachylepis) from Angola and the Democratic Republic of the Congo. , 2019, Zootaxa.

[19]  A. Leaché,et al.  Lifting the blue-headed veil – integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae) , 2018 .

[20]  A. Bauer,et al.  Systematics and phylogeography of the widely distributed African skink Trachylepis varia species complex. , 2018, Molecular phylogenetics and evolution.

[21]  W. Conradie,et al.  Preliminary herpetological survey of Ngonye Falls and surrounding regions in south-western Zambia , 2017 .

[22]  M. Bates,et al.  A review of Cordylus machadoi (Squamata: Cordylidae) in southwestern Angola, with the description of a new species from the Pro-Namib desert. , 2016, Zootaxa.

[23]  Charles W. Linkem,et al.  A hybrid phylogenetic-phylogenomic approach for species tree estimation in African Agama lizards with applications to biogeography, character evolution, and diversification. , 2014, Molecular phylogenetics and evolution.

[24]  M. Bates,et al.  A molecular phylogeny of the African plated lizards, genus Gerrhosaurus Wiegmann, 1828 (Squamata: Gerrhosauridae), with the description of two new genera. , 2013, Zootaxa.

[25]  G. J. Measey,et al.  A molecular phylogeny for sub-Saharan amphisbaenians , 2013 .

[26]  M. Vences,et al.  Molecular phylogeny of African hinge‐back tortoises (Kinixys): implications for phylogeography and taxonomy (Testudines: Testudinidae) , 2012 .

[27]  D. Broadley A review of the genus Mabuya in southeastern Africa (Sauria: Scincidae) , 2000 .

[28]  N. Jacobsen THE STATUS OF AGAMA ACULEATA ARMATA PETERS 1854, (REPTILIA: AGAMIDAE) , 1992 .

[29]  W. Branch,et al.  A REPORT ON A SMALL COLLECTION OF REPTILES FROM SOUTHERN ANGOLA , 1992 .

[30]  W. D. Haacke New herpetological records from South West Africa , 1970 .

[31]  H. W. Parker The Lizards of South Africa , 1946, Nature.

[32]  W. Conradie,et al.  Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species , 2021, Vertebrate Zoology.

[33]  W. Conradie,et al.  The Amphibians of Angola: Early Studies and the Current State of Knowledge , 2019, Biodiversity of Angola.

[34]  Gabriela B Bittencourt-Silva A herpetological survey of western Zambia , 2019 .

[35]  Transvaal Museum,et al.  THE BURROWING GECKOS OF SOUTHERN AFRICA , 2 ( RE PTILIA : GEKKONIDAE ) , 2009 .

[36]  W. Branch,et al.  The skink Mabuya ivensii: new records from Zambia and Zaire, and the status of the subspecies septemlineata Laurent 1964 and the genus Lubuya Horton 1972 , 1993 .

[37]  D. Broadley The reptiles and amphibians of Zambia , 1971 .

[38]  R. F. Laurent Reptiles et amphibiens de l'Angola (troisième contribution) , 1964 .

[39]  Barbosa du Bocage Herpétologie d'Angola et du Congo : , 1895 .