Hyenas Around The City (Kashan, Iran)

This paper presents a taphonomic study of fauna remains of domestic and wild mammals found in a striped hyaena (Hyaena hyaena) den at Kaftar Khoun in the Karkars Piedmont near the city gate of Kashan (Iran). The Kaftar Khoun faunal assemblage is characterized by a low degree of bone breakage with many of the long bones complete, an intermediate frequency of tooth marking and a moderate amount of weathering damage to the bones. The species list, and mortality profiles of the main taxa, suggests that the hyenas collected remains of domestic stock that died naturally or were hunted/scavenged (e. g. mules, donkeys), while the canids represent prey killed during conflicts over carcasses or were scavenged from road kills. The Kaftar Khoun den offers insights into the behaviour of striped hyenas in peri-urban environments. It shows that their behavioral adaptations are directly connected to modifications in their environment such that it may be considered as a commensal animal associated with human activities.

[1]  E. Trinkaus,et al.  Carnivores and their prey in the Wezmeh Cave (Kermanshah, Iran): a Late Pleistocene refuge in the Zagros , 2009 .

[2]  J. Bahain,et al.  Neandertals paleoenvironment in Western Provence: The contribution of Les Auzières 2 (Méthamis, Vaucluse, France) , 2009 .

[3]  D. Todisco,et al.  Bone Weathering in a Periglacial Environment: The Tayara Site (KbFk-7), Qikirtaq Island, Nunavik (Canada) , 2009 .

[4]  H. Monchot Des hyènes tachetées au Pléistocène supérieur dans le Zagros (grotte Wezmeh, Iran) , 2008 .

[5]  C. Egeland,et al.  Taphonomic Analysis of a Modern Spotted Hyena (Crocuta crocuta) Den from Nairobi, Kenya , 2008 .

[6]  M. Prendergast,et al.  Taphonomic Analyses of a Hyena Den and a Natural-Death Assemblage Near Lake Eyasi (Tanzania) , 2008 .

[7]  J. Peterhans,et al.  Spotted hyena (Crocuta crocuta) den use and taphonomy in the Masai Mara National Reserve, Kenya , 2007 .

[8]  C. Diedrich,et al.  Prey deposits and den sites of the Upper Pleistocene hyena Crocuta crocuta spelaea (Goldfuss, 1823) in horizontal and vertical caves of the Bohemian Karst (Czech Republic) , 2006 .

[9]  M. Colbert The facial skeleton of the early Oligocene colodon (perissodactyla, tapiroidea) , 2005 .

[10]  R. Lacruz,et al.  Bone Accumulations at Brown Hyena (Parahyaena brunnea) Den Sites in the Makgadikgadi Pans, Northern Botswana: Taphonomic, Behavioral and Palaeoecological Implications , 2005 .

[11]  H. Monchot Un assemblage original au Paléolithique moyen : le repaire à hyènes, porcs-épics et hominidés de la grotte Geula (Mont Carmel, Israël) , 2005 .

[12]  M. Stiner Comparative ecology and taphonomy of spotted hyenas, humans, and wolves in Pleistocene Italy , 2004 .

[13]  T. Pickering Reconsideration of criteria for differentiating faunal assemblages accumulated by hyenas and hominids , 2002 .

[14]  H. Monchot,et al.  Représentation squelettique au Paléolithique inférieur, le site d'Holon (Israël) , 2002 .

[15]  E. Lotan Feeding the scavengers. Actualistic taphonomy in the Jordan Valley, Israel , 2000 .

[16]  J. Frembgen The magicality of the hyena : Beliefs and practices in West and South Asia , 1998 .

[17]  J. Brugal,et al.  Les repaires d'hyènes des cavernes en Europe occidentale : présentation et comparaison de quelques assemblages osseux. , 1997 .

[18]  Jean-Philip Brugal,et al.  Comparative study of bone assemblages made by recent and Pleistocene Hyenids. , 1997 .

[19]  P. Fossé Variabilité des assemblages osseux créés par l'Hyène des Cavernes , 1997 .

[20]  M. Qumsiyeh Mammals of the Holy Land , 1996 .

[21]  K. Lupo Hadza Bone Assemblages and Hyena Attrition: An Ethnographic Example of the Influence of Cooking and Mode of Discard on the Intensity of Scavenger Ravaging , 1995 .

[22]  P. Fossé Le rôle de l'hyène dans la formation des associations osseuses : 150 ans de controverses [L'apport des anciens textes de préhistoire et de paléontologie du Quaternaire aux études taphonomiques actuelles] , 1995 .

[23]  Y. Lam Variability in the behaviour of spotted hyaenas as taphonomic agents , 1992 .

[24]  A. Stuart MAMMALIAN EXTINCTIONS IN THE LATE PLEISTOCENE OF NORTHERN EURASIA AND NORTH AMERICA , 1991, Biological reviews of the Cambridge Philosophical Society.

[25]  K. Cruz-Uribe Distinguishing Hyena from Hominid Bone Accumulations , 1991 .

[26]  Patricia Smith,et al.  The effects of striped hyaena activity on human remains , 1988 .

[27]  Lewis R. Binford,et al.  Hyena scavenging behavior and its implications for the interpretation of faunal assemblages from FLK 22 (the zinj floor) at olduvai gorge , 1988 .

[28]  L. Todd,et al.  Long bone fragmentation and interpretation of faunal assemblages: Approaches to comparative analysis , 1988 .

[29]  D. Boneh Mystical Powers of Hyenas:Interpreting a Bedouin Belief , 1987 .

[30]  Badoux Dm,et al.  Descriptive and functional morphology of the neck and forelimb of the striped hyena (Hyaena hyaena, L. 1758). , 1986 .

[31]  C. K. Brain The Hunters or the Hunted?: An Introduction to African Cave Taphonomy , 1983, Geological Magazine.

[32]  L. Binford Bones: Ancient Men and Modern Myths , 1981 .

[33]  G. Haynes Evidence of carnivore gnawing on Pleistocene and Recent mammalian bones , 1980, Paleobiology.

[34]  P. Richardson Carnivore damage to antelope bones and its archaeological implications , 1980 .

[35]  J. Henschel,et al.  Implications of a Spotted Hyaena Bone Assemblage in the Namib Desert , 1979 .

[36]  V. Eisenmann Les métapodes d'Equus sensu lato (Mammalia, Périssodactyla) , 1979 .

[37]  Anna K. Behrensmeyer,et al.  Taphonomic and ecologic information from bone weathering , 1978, Paleobiology.

[38]  T. Roberts The mammals of Pakistan , 1977 .

[39]  H. Kruuk Feeding and social behaviour of the striped hyaena (Hyaena vulgaris Desmarest) , 1976 .

[40]  G. J. Miller A Study of Cuts, Grooves, and Other Marks on Recent and Fossil Bone: II Weathering Cracks, Fractures, Splinters, and Other Similar Natural Phenomena , 1975 .

[41]  A. J. Sutcliffe,et al.  Spotted Hyaena: Crusher, Gnawer, Digester and Collector of Bones , 1970, Nature.

[42]  D. Harrison,et al.  The mammals of Arabia , 1964 .

[43]  G. Simpson Large Pleistocene felines of North America. American Museum novitates ; no. 1136 , 1941 .