PRELIMINARY RESULTS OF A PHYLOGENETIC ANALYSIS OF THE PTEROSAURS FROM WESTERN LIAONING AND SURROUNDING AREAS

Many pterosaurs were found recently in western Liaoning and the surrounding areas. Of the 17 genera and 18 species named, only four genera can be assigned confidently to known families. Due to the incompleteness of pterosaur remains (some are represented by postcranial material, whereas others are known from skulls only), the phylogenetic relationships of these pterosaurs are unclear. Based on the modified character matrix of Kellner (80 characters), the relationships of Liaoning pterosaurs (56 taxa, including outgroups) are explored. Preliminary results show that: 1) Dendrorhynchoides and Jeholopterus belong to Anurognathidae; 2) the clade formed by Boreopterus and Feilongus is basal to other ornithocheirids; 3) Liaoningopterus is a primitive form of Anhangueridae; 4) Beipiaopterusis a possible basal ctenochasmatoid; 5) Eosipterus is basal to Germanodacylidae; 6) Eoazhdarcho and Eopteranodon may be early forms of Azhdarchoidea; 7) Chaoyangopterus and Jidapterus may belong to Azhdarchoidea, but they are more derived than Eopteranodon and Eoazhdarcho, and 8) three Chinese tapejarids form a monophyletic group.

[1]  Y. Tomida,et al.  A partial skeleton of titanosaurian sauropod from the Early Cretaceous of Toba City, central Japan , 2006 .

[2]  Zhonghe Zhou,et al.  Pterosaur diversity and faunal turnover in Cretaceous terrestrial ecosystems in China , 2005, Nature.

[3]  Lü Junchang,et al.  New Tapejarid Pterosaur from Western Liaoning, China , 2005 .

[4]  Ji Qiang,et al.  A New Ornithocheirid from the Early Cretaceous of Liaoning Province, China , 2005 .

[5]  Zhonghe Zhou,et al.  A new pterosaur (Pterodactyloidea, Tapejaridae) from the Early Cretaceous Jiufotang Formation of western Liaoning, China and its implications for biostratigraphy , 2003 .

[6]  Zhonghe Zhou,et al.  A nearly completely articulated rhamphorhynchoid pterosaur with exceptionally well-preserved wing membranes and “hairs” from Inner Mongolia, northeast China , 2002 .

[7]  Q. Ji,et al.  Biostratigraphy of new pterosaurs from China , 1999, Nature.

[8]  A. Kellner,et al.  Cranial remains of Quetzalcoatlus (Pterosauria, Azhdarchidae) from Late Cretaceous sediments of Big Bend National Park, Texas , 1996 .

[9]  S. Bennett Taxonomy and systematics of the Late Cretaceous pterosaur Pteranodon (Pterosauria, Pterodactyloidea) , 1994 .

[10]  Wayne P. Maddison,et al.  Macclade: Analysis of Phylogeny and Character Evolution/Version 3 , 1992 .

[11]  S. Bennett A pteranodontid pterosaur from the Early Cretaceous of Peru, with comments on the relationships of Cretaceous pterosaurs , 1989, Journal of Paleontology.

[12]  S. Howse,et al.  On the cervical vertebrae of the Pterodactyloidea (Reptilia: Archosauria) , 1986 .

[13]  Jiqiang,et al.  New Azhdarchid Pterosaur from the Early Cretaceous of Western Liaoning , 2005 .

[14]  Zhang Baokun New Pterodactyl oid Pterosaur from the Yixian Formation of Western Liaoning , 2005 .

[15]  DONGZhiming,et al.  A New Ctenochasmatid Pterosaur from the Early Cretaceous of Liaoning Province , 2005 .

[16]  A. Kellner New information on the Tapejaridae (Pterosauria, Pterodactyloidea) and discussion of the relationships of this clade , 2004 .

[17]  D. Unwin On the phylogeny and evolutionary history of pterosaurs , 2003, Geological Society, London, Special Publications.

[18]  Lü Junchang A NEW PTEROSAUR: BEIPIAOPTERUS CHENIANUS, GEN. ET SP. NOV. (REPTILIA: PTEROSAURIA) FROM WESTERN LIAONING PROVINCE OF CHINA , 2003 .

[19]  Li Jian A NEW LOWER CRETACEOUS SINOPTERID PTEROSAUR FROM THE WESTERN LIAONING,CHINA , 2003 .

[20]  W. Xiao TWO NEW PTERODACTYLOID PTEROSAURS FROM THE EARLY CRETACEOUS JIUFOTANG FORMATION OF WESTERN LIAONING,CHINA , 2003 .

[21]  A. Kellner Pterosaur phylogeny and comments on the evolutionary history of the group , 2003, Geological Society, London, Special Publications.

[22]  Lü Junchang SOFT TISSUE IN AN EARLY CRETACEOUS PTEROSAUR FROM LIAONING PROVINCE, CHINA , 2002 .

[23]  D. Unwin,et al.  On the systematic and stratigraphic significance of pterosaurs from the Lower Cretaceous Yixian Formation (Jehol Group) of Liaoning, China , 2000 .

[24]  D. Unwin The phylogeny of the Pterosauria , 1992 .