Breathing device of a new Streptaulus species from Vietnam extends understanding of the function and structure of respiratory tubes in cyclophoroids (Gastropoda: Caenogastropoda Pupinidae)

The operculum of the terrestrial caenogastropod superfamily Cyclophoroidea tightly seals the aperture when the animal with- draws the entire soft body into the shell. For at least most species the operculum presents a barrier to gas exchange. Many cyclo- phoroideans have resolved this problem by evolving shell devices that provide alternative routes for gas exchange. These modi fi ca-tions range from a simple slit- or groove-like channel to a snorkel- like tube at the peristome (Rees, 1964). Elaborate examples are found among Asian members of the family Alycaeidae (Godwin-Austen, 1886; Páll-Gergely, Naggs & Asami, 2016). Their sutural tube has no direct external opening. Instead, many microtunnels extend down from the sutural tube towards their opening near the umbilicus of the body whorl. The sutural tube possesses a pore that opens internally behind the point at which the operculum seals the aperture, allowing gas exchange between the body

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