An Ecosystem Management Strategy for Sierran Mixed-Conifer Forests

Current Sierra Nevada forest management is often focused on strategically reducing fuels without an explicit strategy for ecological restoration across the landscape matrix. Summarizing recent scientific literature, we suggest managers produce different stand structures and densities across the landscape using topographic variables (i.e., slope shape, aspect, and slope position) as a guide for varying treatments. Local cool or moist areas, where historically fire would have burned less frequently or at lower severity, would have higher density and canopy cover, providing habitat for sensitive species. In contrast upper, southern-aspect slopes would have low densities of large fire-resistant trees. For thinning, marking rules would be based on crown strata or age cohorts and species, rather than uniform diameter limits. Collectively, our management recommendations emphasize the ecological role of fire, changing climate conditions, sensitive wildlife habitat, and the importance of forest structure heterogeneity.

[1]  S. Stephens Fuel loads, snag abundance, and snag recruitment in an unmanaged Jeffrey pine–mixed conifer forest in Northwestern Mexico , 2004 .

[2]  James K. Agee KEYNOTE ADDRESS: The Role of Silviculture in Restoring Fire-Adapted Ecosystems , 2007 .

[3]  J. Negrón,et al.  The effectiveness of vegetation management practices for prevention and control of bark beetle infestations in coniferous forests of the western and southern United States , 2007 .

[4]  W. Zielinski,et al.  HOME RANGE CHARACTERISTICS OF FISHERSIN CALIFORNIA , 2004 .

[5]  Christopher R. Keyes,et al.  Quantifying Stand Targets for Silvicultural Prevention of Crown Fires , 2002 .

[6]  K. O’Hara,et al.  Stocking Control Procedures for Multiaged Ponderosa Pine Stands in the Inland Northwest , 2003 .

[7]  D. Urban,et al.  Interactions between forest heterogeneity and surface fire regimes in the southern Sierra Nevada , 1999 .

[8]  J. Battles,et al.  Spatial elements of mortality risk in old-growth forests. , 2008, Ecology.

[9]  S. Stephens,et al.  Thinning, burning, and thin-burn fuel treatment effects on soil properties in a Sierra Nevada mixed-conifer forest , 2007 .

[10]  Bruce R. Hartsough,et al.  The economics of alternative fuel reduction treatments in western United States dry forests: Financial and policy implications from the National Fire and Fire Surrogate Study , 2008 .

[11]  Malcolm P. North,et al.  Initial response of a mixed-conifer understory plant community to burning and thinning restoration treatments , 2007 .

[12]  Jiquan Chen,et al.  Soil respiration response to experimental disturbances over 3 years , 2006 .

[13]  P. Hessburg,et al.  Re-examining fire severity relations in pre-management era mixed conifer forests: inferences from landscape patterns of forest structure , 2007, Landscape Ecology.

[14]  Jan W. van Wagtendonk,et al.  Fire as an Ecological Process , 2006 .

[15]  S. Stephens,et al.  Prescribed fire, soils, and stream water chemistry in a watershed in the Lake Tahoe Basin, California , 2004 .

[16]  Paul F. Hessburg,et al.  Fire and aquatic ecosystems of the western USA: current knowledge and key questions , 2003 .

[17]  Carl N. Skinner,et al.  Basic principles of forest fuel reduction treatments , 2005 .

[18]  F. H. Tainter,et al.  Principles of Forest Pathology , 1996 .

[19]  B. Wing,et al.  Stability of the large tree component in treated and untreated late-seral interior ponderosa pine stands , 2008 .

[20]  M. North,et al.  Mixed‐conifer understory response to climate change, nitrogen, and fire , 2008 .

[21]  Harold S. J. Zald,et al.  Initial tree regeneration responses to fire and thinning treatments in a Sierra Nevada mixed-conifer forest, USA , 2008 .

[22]  M. Barbour,et al.  Influence of Fire and El Niño on Tree Recruitment Varies by Species in Sierran Mixed Conifer , 2005, Forest Science.

[23]  Anne E. Black,et al.  Cross-Scale Analysis of Fire Regimes , 2007, Ecosystems.

[24]  W. McShea THE INFLUENCE OF ACORN CROPS ON ANNUAL VARIATION IN RODENT AND BIRD POPULATIONS , 2000 .

[25]  A. Taylor IDENTIFYING FOREST REFERENCE CONDITIONS ON EARLY CUT‐OVER LANDS, LAKE TAHOE BASIN, USA , 2004 .

[26]  A. B. Carey Biocomplexity and restoration of biodiversity in temperate coniferous forest: inducing spatial heterogeneity with variable‐density thinning , 2003 .

[27]  S. Jackson,et al.  Novel climates, no‐analog communities, and ecological surprises , 2007 .

[28]  Scott L. Stephens,et al.  Experimental fuel treatment impacts on forest structure, potential fire behavior, and predicted tree mortality in a California mixed conifer forest , 2005 .

[29]  James K. Brown Handbook for inventorying downed woody material , 1974 .

[30]  S. Stephens,et al.  FEDERAL FOREST‐FIRE POLICY IN THE UNITED STATES , 2005 .

[31]  Michael A. Crimmins,et al.  Synoptic climatology of extreme fire‐weather conditions across the southwest United States , 2006 .

[32]  R. Arkley Soil Moisture Use by Mixed Conifer Forest in a Summer‐Dry Climate , 1981 .

[33]  Robert F. Powers,et al.  Restoring Fire-Adapted Ecosystems: Proceedings of the 2005 National Silviculture Workshop , 2007 .

[34]  T. Swetnam,et al.  Warming and Earlier Spring Increase Western U.S. Forest Wildfire Activity , 2006, Science.

[35]  Robert J. Naiman,et al.  Fire in the Riparian Zone: Characteristics and Ecological Consequences , 2007, Ecosystems.

[36]  J. Franklin,et al.  Facilitative and competitive effects of a N-fixing shrub on white fir saplings , 2006 .

[37]  A. Taylor,et al.  Fire and persistence of montane chaparral in mixed conifer forest landscapes in the northern Sierra Nevada, Lake Tahoe Basin, California, USA1 , 2005 .

[38]  Kathleen A. Dwire,et al.  Fire and riparian ecosystems in landscapes of the western USA , 2003 .

[39]  Thomas C. Peterson,et al.  Maximum and Minimum Temperature Trends for the Globe , 1997 .

[40]  Robin J. Innes,et al.  Habitat Associations of Dusky-footed Woodrats (Neotoma fuscipes) in Mixed-conifer Forest of the Northern Sierra Nevada , 2007 .

[41]  A. Gray,et al.  Influence of light and soil moisture on Sierran mixed-conifer understory communities , 2005, Plant Ecology.

[42]  S. Stephens,et al.  Spatial patterns of large natural fires in Sierra Nevada wilderness areas , 2007, Landscape Ecology.

[43]  N. Stephenson,et al.  Apparent climatically induced increase of tree mortality rates in a temperate forest. , 2007, Ecology letters.

[44]  Kevin S. McKelvey,et al.  The California Spotted Owl: A Technical Assessment of Its Current Status. , 1992 .

[45]  Andrew B. Carey,et al.  Wildlife and vegetation of unmanaged Douglas-Fir forests , 1991 .

[46]  J W Thomas,et al.  Guidelines for maintaining and enhancing wildlife habitat in forest management in the blue mountains of Oregon and Washington. , 1977 .

[47]  K. O’Hara,et al.  Masam – a flexible stand density management model for meeting diverse structural objectives in multiaged stands , 1999 .

[48]  T. M. Bonnicksen,et al.  Reconstrution of a Presettlement Giant Sequoia‐Mixed Conifer Forest Community Using the Aggregation Approach , 1982 .

[49]  M. Morrison,et al.  Foraging Behavior of Bark-Foraging Birds in the Sierra Nevada , 1987 .

[50]  M. Finney Design of Regular Landscape Fuel Treatment Patterns for Modifying Fire Growth and Behavior , 2001, Forest Science.

[51]  Sierra Nevada Ecosystem,et al.  Sierra Nevada Ecosystem Project final report to Congress , 1997 .

[52]  Malcolm P. North,et al.  Fuel treatment effects on tree‐based forest carbon storage and emissions under modeled wildfire scenarios , 2009 .

[53]  P. Hessburg,et al.  Dry forests and wildland fires of the inland Northwest USA: Contrasting the landscape ecology of the pre-settlement and modern eras , 2005 .

[54]  D. Rizzo,et al.  Present and past old-growth forests of the Lake Tahoe Basin, Sierra Nevada, US , 2002 .

[55]  E. Forsman,et al.  Diets and foraging behavior of northern Spotted Owls in Oregon , 2004 .

[56]  Malcolm P. North,et al.  Forest Stand Structure and Pattern of Old-Growth Western Hemlock/Douglas-Fir and Mixed-Conifer Forests , 2004, Forest Science.

[57]  S. Stephens,et al.  Coarse woody debris and canopy cover in an old-growth Jeffrey pine-mixed conifer forest from the Sierra San Pedro Martir, Mexico , 2007 .

[58]  R. Neilson,et al.  Climate change effects on vegetation distribution, carbon, and fire in California , 2003 .

[59]  Neil G. Sugihara,et al.  Fire in California's ecosystems , 2006 .

[60]  M. North Restoring forest health: fire and thinning effects on mixed-conifer forests , 2006 .

[61]  H. Erickson,et al.  Effects of vegetation patches on soil nutrient pools and fluxes within a mixed-conifer forest , 2005 .

[62]  D. Falk Discovering the Future, Creating the Past: Some Reflections on Restoration , 1990, Restoration & Management Notes.

[63]  Jim InnesJ. Innes,et al.  Comparison of thinning and prescribed fire restoration treatments to Sierran mixed-conifer historic conditions , 2007 .

[64]  R. Everett,et al.  Continuity in fire disturbance between riparian and adjacent sideslope Douglas-fir forests , 2003 .

[65]  Jerry F. Franklin,et al.  Creating a forestry for the 21st century : the science of ecosystem management , 1997 .

[66]  Michael R. Wagner,et al.  Concepts of forest health: Utilitarian and ecosystem perspectives , 1994 .

[67]  S. Stephens,et al.  Effects of prescribed fire on a Sierra Nevada (California, USA) stream and its riparian zone , 2005 .

[68]  S. Stephens,et al.  Comparing modern and past fire regimes to assess changes in prehistoric lightning and anthropogenic ignitions in a Jeffrey pine – mixed conifer forest in the Sierra San Pedro Mártir, Mexico , 2007 .

[69]  S. Stephens,et al.  Fire and fire surrogate treatment effects on leaf litter arthropods in a western Sierra Nevada mixed-conifer forest , 2006 .

[70]  Scott L. Stephens,et al.  Fire Climbing in the Forest: A Semiqualitative, Semiquantitative Approach to Assessing Ladder Fuel Hazards , 2007 .

[71]  J. Tappeiner,et al.  Response of old-growth conifers to reduction in stand density in western Oregon forests. , 2002, Tree physiology.

[72]  Scott L. Stephens,et al.  Prehistoric fire area and emissions from California's forests, woodlands, shrublands, and grasslands , 2007 .

[73]  M. North,et al.  Microhabitat Associations of Northern Flying Squirrels in Burned and Thinned Forest Stands of the Sierra Nevada , 2007 .

[74]  A. Taylor,et al.  Spatial and temporal variation of fire regimes in a mixed conifer forest landscape, Southern Cascades, California, USA , 2001 .

[75]  E. Knapp,et al.  Heterogeneity in fire severity within early season and late season prescribed burns in a mixed-conifer forest* , 2006 .

[76]  W. Zielinski,et al.  RESTING HABITAT SELECTION BY FISHERS IN CALIFORNIA , 2004 .

[77]  R. Barrett,et al.  Foraging and Habitat Relationships of Insect-Gleaning Birds in a Sierra Nevada Mixed-Conifer Forest , 1985 .

[78]  Jason J. Moghaddas,et al.  Tree mortality patterns following prescribed fires in a mixed conifer forest , 2006 .

[79]  M. North,et al.  Effect of thinning and prescribed fire restoration treatments on woody debris and snag dynamics in a Sierran old-growth, mixed-conifer forest , 2006 .

[80]  E. C. Farmer,et al.  DIET OF FISHERS (MARTES PENNANTI) AT THE SOUTHERNMOST EXTENT OF THEIR RANGE , 1999 .

[81]  M. Finney,et al.  Modeling wildfire risk to northern spotted owl (Strix occidentalis caurina) habitat in Central Oregon, USA , 2007 .

[82]  T. Dawson,et al.  Conifer Seedling Survival Under Closed-Canopy And Manzanita Patches In the Sierra Nevada , 2008 .

[83]  Malcolm P. North,et al.  Initial tree mortality and insect and pathogen response to fire and thinning restoration treatments in an old-growth mixed-conifer forest of the Sierra Nevada, California , 2008 .

[84]  D. Lindenmayer,et al.  Conserving Forest Biodiversity: A Comprehensive, Multiscaled Approach , 2002 .

[85]  George R. Hess,et al.  Communicating clearly about conservation corridors , 2001 .

[86]  M. North,et al.  Spatial and Temporal Variability of Canopy Microclimate in a Sierra Nevada Riparian Forest , 2008 .

[87]  C. Skinner A tree-ring based fire history of riparian reserves in the Klamath Mountains. , 2003 .

[88]  C. Allen,et al.  ECOLOGICAL RESTORATION OF SOUTHWESTERN PONDEROSA PINE ECOSYSTEMS: A BROAD PERSPECTIVE , 2002 .

[89]  S. Stephens,et al.  Forest structure and mortality in an old-growth Jeffrey pine-mixed conifer forest in north-western Mexico , 2005 .

[90]  J. R. Waters,et al.  General biology of major prey species of the California spotted owl , 1992 .

[91]  Charles W. McHugh,et al.  Simulation of long-term landscape-level fuel treatment effects on large wildfires , 2006 .

[92]  Klaus H. Barber,et al.  Stewardship and fireshed assessment: a process for designing a landscape fuel treatment strategy. , 2007 .

[93]  Jerry F. Franklin,et al.  Spatial Aspects of Structural Complexity in Old-Growth Forests , 2004, Journal of Forestry.

[94]  William J. Zielinski,et al.  Individual legacy trees influence vertebrate wildlife diversity in commercial forests , 2004 .

[95]  K. O’Hara Dynamics and Stocking-Level Relationships of Multi-Aged Ponderosa Pine Stands , 1996 .

[96]  M. North,et al.  Truffle abundance in riparian and upland mixed-conifer forest of California's southern Sierra Nevada , 2005 .

[97]  J. Wiedemann My First Summer in the Sierra , 2007 .

[98]  Norman L. Miller,et al.  POTENTIAL IMPACTS OF CLIMATE CHANGE ON CALIFORNIA HYDROLOGY 1 , 2003 .

[99]  E. Knapp,et al.  Role of burning season on initial understory vegetation response to prescribed fire in a mixed conifer forest , 2007 .

[100]  S. Stephens,et al.  Climate change and forests of the future: managing in the face of uncertainty. , 2007, Ecological applications : a publication of the Ecological Society of America.

[101]  K. O’Hara SILVICULTURE FOR STRUCTURAL DIVERSITY : A NEW LOOK AT MULTIAGED SYSTEMS , 1998 .

[102]  S. Stephens,et al.  Wildfire and Spatial Patterns in Forests in Northwestern Mexico: The United States Wishes It Had Similar Fire Problems , 2008 .

[103]  Jason J. Moghaddas,et al.  The National Fire and Fire Surrogate study: effects of fuel reduction methods on forest vegetation structure and fuels. , 2009, Ecological applications : a publication of the Ecological Society of America.

[104]  T. Kolb,et al.  Tree Size and Drought Affect Ponderosa Pine Physiological Response to Thinning and Burning Treatments , 2004, Forest Science.

[105]  Maggi Kelly,et al.  Interactions Among Wildland Fires in a Long-Established Sierra Nevada Natural Fire Area , 2009, Ecosystems.

[106]  Harold S. J. Zald,et al.  Influence of soil thickness on stand characteristics in a Sierra Nevada mixed-conifer forest , 2007, Plant and Soil.

[107]  Joseph W. Sherlock Integrating stand density management with fuel reduction , 2007 .

[108]  S. Schneider,et al.  Emissions pathways, climate change, and impacts on California. , 2004, Proceedings of the National Academy of Sciences of the United States of America.

[109]  J. D. Johnston,et al.  Historical perspectives on forests of the Sierra Nevada and the transverse ranges of southern California; forest conditions at the turn of the century , 1992 .

[110]  S. Stephens,et al.  Spatial distribution of regeneration patches in an old-growth Pinus jeffreyi-mixed conifer forest in northwestern Mexico , 2005 .

[111]  M. Hurteau,et al.  Fire suppression and fuels treatment effects on mixed-conifer carbon stocks and emissions. , 2009, Ecological applications : a publication of the Ecological Society of America.

[112]  L. Tevis Autumn Foods of Chipmunks and Golden-Mantled Ground Squirrels in the Northern Sierra Nevada , 1952 .

[113]  R. Graham,et al.  Roles of weathered bedrock and soil in seasonal water relations of Pinus Jeffreyi and Arctostaphylos patula , 2001 .

[114]  W. Keeton,et al.  Emulating Natural Disturbance Regimes: an Emerging Approach for Sustainable Forest Management , 2008 .

[115]  N. Stephenson REFERENCE CONDITIONS FOR GIANT SEQUOIA FOREST RESTORATION: STRUCTURE, PROCESS, AND PRECISION , 1999 .

[116]  Peter Z. Fulé,et al.  Restoring Ecosystem Health in Ponderosa Pine Forests of the Southwest , 1997, Journal of Forestry.

[117]  D. M. Smith,et al.  The Practice Of Silviculture: Applied Forest Ecology , 2014 .

[118]  J. B. Leiberg Forest Conditions in the Northern Sierra Nevada, California , 2009 .

[119]  C. Skinner,et al.  Probability of tree survival after wildfire in an interior pine forest of northern California: Effects of thinning and prescribed fire , 2007 .

[120]  J. W. Wagtendonk,et al.  An Overview of Fire in the Sierra Nevada , 1996 .

[121]  Seth Bigelow,et al.  HABITAT ASSOCIATIONS OF SMALL MAMMALS AT TWO SPATIAL SCALES IN THE NORTHERN SIERRA NEVADA , 2006 .

[122]  D. Falk Process-centred restoration in a fire-adapted ponderosa pine forest , 2006 .

[123]  M. North,et al.  Association of weather and nest-site structure with reproductive success in California spotted owls , 2000 .

[124]  J. Battles,et al.  Growth Response of Mature Trees versus Seedlings to Gaps Associated with Group Selection Management in the Sierra Nevada, California , 2008 .

[125]  J. Rappole,et al.  The Science of Conservation Planning: Habitat Conservation Under The Endangered Species Act , 1997 .

[126]  M. North,et al.  Canopy microclimate response to pattern and density of thinning in a Sierra Nevada forest , 2009 .

[127]  J. Franklin,et al.  The effects of fire on soil nitrogen associated with patches of the actinorhizal shrub Ceanothus cordulatus , 2003, Plant and Soil.

[128]  C. Millar,et al.  THE ROLE OF CLIMATE CHANGE IN INTERPRETING HISTORICAL VARIABILITY , 1999 .

[129]  G. Mroz,et al.  Microclimate in Forest Ecosystem and Landscape Ecology , 1999 .

[130]  A. Taylor,et al.  SPATIAL PATTERNS AND CONTROLS ON HISTORICAL FIRE REGIMES AND FOREST STRUCTURE IN THE KLAMATH MOUNTAINS , 2003 .

[131]  Robert F. Powers,et al.  The effects of wildfire, salvage logging, and post-fire N-fixation on the nutrient budgets of a Sierran forest , 2005 .

[132]  C. Skinner Silviculture and forest management under a rapidly changing climate , 2007 .

[133]  N. McDowell,et al.  Perpetuating old ponderosa pine , 2007 .

[134]  D. Rizzo,et al.  Patterns of Mortality in an Old-Growth Mixed-Conifer Forest of the Southern Sierra Nevada, California , 2005, Forest Science.

[135]  Jeffrey R Dunk,et al.  Using forest inventory data to assess fisher resting habitat suitability in California. , 2006, Ecological applications : a publication of the Ecological Society of America.

[136]  W. W. Miller,et al.  PRESCRIBED FIRE EFFECTS ON FOREST FLOOR AND SOIL NUTRIENTS IN A SIERRA NEVADA FOREST , 2006 .

[137]  R. Schmid,et al.  Fire in Sierra Nevada Forests: A Photographic Interpretation of Ecological Change Since 1849 , 2001 .

[138]  Jerry F. Franklin,et al.  The Structure of Natural Young , Mature , and Old-Growth Douglas-Fir Forests in Oregon and Washington , 2010 .

[139]  M. Barbour,et al.  Sixty Years of Change in Californian Conifer Forests of the San Bernardino Mountains , 1995 .

[140]  Daniel R. Miller,et al.  Effects of post-wildfire erosion on channel environments, Boise River, Idaho , 2003 .

[141]  Malcolm P. North,et al.  Vegetation and ecological characteristics of mixed-conifer and red fir forests at the Teakettle Experimental Forest , 2002 .

[142]  Jay D. Miller,et al.  Quantitative Evidence for Increasing Forest Fire Severity in the Sierra Nevada and Southern Cascade Mountains, California and Nevada, USA , 2009, Ecosystems.

[143]  Malcolm P. North,et al.  NEST TREES OF NORTHERN FLYING SQUIRRELS IN THE SIERRA NEVADA , 2005 .

[144]  Patrick N. Halpin,et al.  Forest gradient response in Sierran landscapes: the physical template , 2000, Landscape Ecology.