Systematic Revision of Trilobites from the Middle Ordovician (Darriwilian) Klimoli Formation of the Zhuozishan Area, Inner Mongolia, China

New morphologic information permits systematic revision of trilobites from the Middle Ordovician Klimoli Formation of the Zhuozishan area, Ordos Basin, Inner Mongolia. The new assemblage is composed of 10 species of the Raphiophoridae, Nileidae, Asaphidae, and Telephinidae. An asaphid, aff. Mioptychopyge lashachungensis (previously Paraptychopyge lashachungensis) displays an intermediate morphology between the Chinese nobiliasaphine genera Mioptychopyge and Zhenganites. The pygidial doublure is regarded as the most significant character to differentiate Symphysurus klimoliensis (previously Nileus klimoliensis) of the Nileidae from such closely allied taxa as Poronileus. A nileid, cf. Peraspis kujandensis displays typical nileid morphology, unlike the type species, Peraspis lineolata, which might turn out to be an asaphid. Ampyx gongwusuensis sp. nov. of the Raphiophoridae is the first record of the genus in the Zhuozishan area and reveals morphologic details that might be employed to resolve Ampyx taxonomy in China. Morphologic differences between A. gongwusuensis and Abulbaspis ordosensis might represent a case of sexual dimorphism.

[1]  Sun Xiao,et al.  Structural Evolution and Hydrocarbon Potential of the Upper Paleozoic Northern Ordos Basin, North China , 2015 .

[2]  Yuandong Zhang,et al.  On the integration of Ordovician conodont and graptolite biostratigraphy: new examples from Gansu and Inner Mongolia in China , 2013 .

[3]  H. B. Whittington Silicified Middle Ordovician Trilobites: Remopleurididae, Trinucleidae, Raphiophoridae, Endymioniidae , 2013 .

[4]  Chen Xu,et al.  Ordovician paleogeography and tectonics of the major paleoplates of China , 2010 .

[5]  L. Popov,et al.  Middle Ordovician (Late Darriwilian) Trilobites from the Northern Betpak-Dala Desert, Central Kazakhstan , 2009 .

[6]  Estes,et al.  Bulletin of the Museum of Comparative Zoology at Harvard College , 2009 .

[7]  N. Hughes Trilobites , 2008, Current Biology.

[8]  Zhou Zhiyi,et al.  Trilobite record of China , 2008 .

[9]  S. Turvey ASAPHOID TRILOBITES FROM THE ARENIG–LLANVIRN OF THE SOUTH CHINA PLATE , 2007 .

[10]  J. R. Ebbestad Trilobites of the Tremadoc Bjørkåsholmen Formation in the Oslo Region, Norway , 1999, Fossils and Strata.

[11]  Zhou Zhi TWO NEW ORDOVICIAN TRILOBITE GENERA FROM WESTERN MARGINAL AREAS OF THE NORTH CHINA PLATFORM , 2006 .

[12]  H. B. Whittington The trilobite family Nileidae: morphology and classification , 2003 .

[13]  H. B. Whittington STYGINA, EOBRONTEUS (ORDOVICIAN STYGINIDAE, TRILOBITA): MORPHOLOGY, CLASSIFICATION, AND AFFINITIES OF ILLAENIDAE , 2000 .

[14]  Yue Li,et al.  Restudy on the trilobites (agnostoids and other polymerids) from the Miaopo Formation (Upper Ordovician) in eastern Yangtze gorge area, Western Hubei , 2000 .

[15]  D. Choi,et al.  Ontogenetic changes of bacculae in Korean asaphid trilobites and their taxonomic implications , 1999, Journal of Paleontology.

[16]  Zy Y. Zhou,et al.  Ordovician trilobites from the Dawangou Formation, Kalpin, Xinjiang, north-west China , 1998 .

[17]  R. Fortey,et al.  Brood pouches in trilobites , 1998, Journal of Paleontology.

[18]  S. Westrop Trilobite systematics, biostratigraphy and palaeoecology of the lower ordovician komstad limestone and huk formations, southern Scandinavia , 1997 .

[19]  D. Choi,et al.  Reappraisal of the Middle Ordovician Trilobites from the Jigunsan Formation, Korea , 1992 .

[20]  R. Fortey,et al.  Classification of the trilobite suborder Asaphina , 1988 .

[21]  R. Fortey The type species of the ordovician trilobitesymphysurus: systematics, functional morphology and terrace ridges , 1986 .

[22]  Zhou Zhiyi,et al.  Ordovician trilobites from North and Northeast China , 1986 .

[23]  Zhiyi. Zhou,et al.  Trilobites from the Ordovician Shihtzupu Formation, Zunyi, Guizhou Province, China , 1984, Transactions of the Royal Society of Edinburgh: Earth Sciences.

[24]  R. Ross,et al.  New Species of Brachiopods and Trilobites from the Middle Ordovician (Whiterock) of southeastern British Columbia , 1978 .

[25]  R. Fortey The Ordovician Trilobites of Spitsbergen. II, Asaphidae, Nileidae, Raphiophoridae and Telephinidae of the Valhallfonna Formation , 1975 .

[26]  W. T. Dean Ordovician trilobites from the Keele Range, northwestern Yukon Territory , 1973 .

[27]  R. Ross Ordovician brachiopods, trilobites, and stratigraphy in eastern and central Nevada , 1970 .

[28]  H. B. Whittington Trilobites of the Ordovician Table Head formationc western Newfoundland , 1965 .

[29]  H. B. Whittington SIXTEEN ORDOVICIAN GENOTYPE TRILOBITES , 1950 .

[30]  Joachim Barrande,et al.  Systême silurien du centre de la Bohême , 1852 .

[31]  Hermann Burmeister Die Organisation der Trilobiten aus ihren lebenden Verwandten entwickelt : nebst einer systematischen Uebersicht aller zeither beschriebenen Arten / von Hermann Burmeister ... Mit 6 Kupfertafeln. , 1843 .

[32]  Hermann Burmeister Die Organisation der Trilobiten, aus ihren lebenden verwandten entwickelt : nebst einer systematischen Uebersicht aller zeither beschriebenen Arten , 1843 .