From simple rules to cycling in community assembly

358.Simulation studies of community assembly have frequently observed two relatedphenomena: (1) the humpty dumpty effect in which communities can not bereconstructed by ‘‘sequential’’ invasions (i.e. single species invasions separated bylong intervals of time) and (2) cycling between sub-communities. To better understandthe mechanisms underlying these phenomena, we analyze a system consisting of twopredators and two prey competing for a shared resource. We show how simpledominance rules (i.e. R

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