An ecophysiological approach to modelling resource fluxes in competing plants

sidering a plant growing in isolation and its response to manipulation of light, CO 2 and N supplies. Secondly, (iv) analyse the extent to which a plant can perceive the competitive plants are introduced and the direct presence of neighbouring plants by signals other than the effects on plant responses in terms of resource deple- quantitative depletion of physical resources; and (v) intetion are considered separately from indirect effects grate resource eVects with non-resource eVects in a more such as potential changes in the quality of resources comprehensive model at the level of the plant stand. Such available (e.g. light quality or soil N sources). In the an ecophysiological approach deals with both intrapast, many studies of plant competition have not specific and inter-specific competition. In developing a established the importance of these indirect effects conceptual model there is no reason to think that changing because they have not established all the processes the neighbouring species should change anything in the involved in competition. This model can be used to model, if the intrinsic capacities of the given species in interpret responses of whole plants to their neighbours modifying the physical resources quantitatively and qualin terms of the relative importance of both the direct itatively are properly described and accounted for at the and indirect effects of competition. scale of the individual plant. The first section of this paper describes a mechanistic,

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