Influence of soil thickness on stand characteristics in a Sierra Nevada mixed-conifer forest

Soil thickness can be an important factor influencing vegetation, yet few spatially-explicit studies have examined soil horizon thickness and vegetation composition in summer-drought forests. We compared seismic and soil penetration measurements of combined A + C and Cr horizon thickness, soil moisture and temperature, and stand variables in a contiguous 4-ha mixed-conifer stand of the Sierra Nevada. Thickness of A + C and Cr horizons were highly variable but were not correlated to each other. Total basal area and canopy cover were positively related with A + C horizon thickness, and shrub cover was positively related with Cr horizon thickness. Basal area of white fir [Abies concolor (Gord and Glend) Lindl.] and incense-cedar [Calocedrus decurrens (Torrey) Florin] were positively correlated with A + C horizon thickness, but there was no relationship between A + C or Cr horizon thickness and basal area of Jeffrey pine (Pinus jeffreyi Grev. and Balf.), sugar pine (P. lambertiana Douglas), or red fir (A. magnifica A. Murray). Both white and red fir seedlings were associated with decreased soil temperature, but only white fir seedlings were positively associated with soil moisture. Soil penetration estimates of soil thickness were similar to seismic estimates for shallow soils (<50 cm depth) but were poorly related on deeper soils. Visual surface conditions and tile probe estimates of soil thickness can be highly misleading because ‘shallow’ areas may have a thick layer of weathered bedrock that can serve as a potential rooting medium for deep-rooted trees and shrubs. In our study only the refraction seismic method had the potential to measure total soil depth that included A + C and Cr horizon thickness.

[1]  J. S. Fralish The Effect of Site Environment on Forest Productivity in the Illinois Shawnee Hills , 1994 .

[2]  Sierra Nevada Ecosystem,et al.  Sierra Nevada Ecosystem Project final report to Congress , 1997 .

[3]  J. Tappeiner,et al.  Effects of Silvicultural Practices and Wildfire on Productivity of Forest Soils , 1997 .

[4]  J. Romanyà Productivity of Pinus radiata plantations in Spain in response to climate and soil , 2004 .

[5]  R. Graham,et al.  Deep regolith: exploring the lower reaches of soil , 2005 .

[6]  Harold S. J. Zald,et al.  Stand Conditions Associated with Tree Regeneration in Sierran Mixed- Conifer Forests , 2005 .

[7]  J. Tappeiner,et al.  Competitive relations between Douglas-fir and Pacific madrone on shallow soils in a Mediterranean climate , 1995 .

[8]  R. Arkley Soil Moisture Use by Mixed Conifer Forest in a Summer‐Dry Climate , 1981 .

[9]  M. Barbour,et al.  Influence of Fire and El Niño on Tree Recruitment Varies by Species in Sierran Mixed Conifer , 2005, Forest Science.

[10]  H. A. Mooney,et al.  Maximum rooting depth of vegetation types at the global scale , 1996, Oecologia.

[11]  T. Bruns Thoughts on the processes that maintain local species diversity of ectomycorrhizal fungi , 1995, Plant and Soil.

[12]  Todd R. Lookingbill,et al.  An empirical approach towards improved spatial estimates of soil moisture for vegetation analysis , 2004, Landscape Ecology.

[13]  Samuel D. Fuhlendorf,et al.  The influence of soil depth on plant species response to grazing within a semi-arid savanna , 1998, Plant Ecology.

[14]  M. Barbour,et al.  Mediterranean climate effects. I. Conifer water use across a Sierra Nevada ecotone. , 2001, American journal of botany.

[15]  P. Reich,et al.  Seed rain, safe sites, competing vegetation, and soil resources spatially structure white pine regeneration and recruitment , 2003 .

[16]  Graham Rc,et al.  Water source utilization by Pinus jeffreyi and Arctostaphylos patula on thin soils over bedrock , 2003 .

[17]  D. Leopold,et al.  Forest community composition and juvenile red spruce (Picea rubens) age-structure and growth patterns in an Adirondack watershed , 1994 .

[18]  D. Markewitz,et al.  How Deep Is Soil?Soil, the zone of the earth's crust that is biologically active, is much deeper than has been thought by many ecologists , 1995 .

[19]  F. P. Haeni Application of seismic refraction methods in groundwater modeling studies in New England , 1986 .

[20]  M. Anderson,et al.  Root distribution and seasonal water status in weathered granitic bedrock under chaparral , 1996 .

[21]  David W. Hosmer,et al.  Applied Logistic Regression , 1991 .

[22]  R. Graham,et al.  Roles of weathered bedrock and soil in seasonal water relations of Pinus Jeffreyi and Arctostaphylos patula , 2001 .

[23]  D. Parker,et al.  Water source utilization by Pinus jeffreyi and Arctostaphylos patula on thin soils over bedrock , 2002, Oecologia.

[24]  R. Graham,et al.  Soil and Weathered Bedrock , 2001 .

[25]  P. J. Fenning,et al.  Comparison of the seismic and ground probing radar methods in geological surveying , 1988 .

[26]  Malcolm P. North,et al.  Vegetation and ecological characteristics of mixed-conifer and red fir forests at the Teakettle Experimental Forest , 2002 .

[27]  T. Spies,et al.  Water content measurement in forest soils and decayed wood using time domain reflectometry , 1995 .

[28]  Luis A. Gallardo,et al.  Evidence for correlation of electrical resistivity and seismic velocity in heterogeneous near‐surface materials , 2003 .

[29]  James A. Doolittle,et al.  Contributions of water supply from the weathered bedrock zone to forest soil quality , 2003 .

[30]  M. Barbour,et al.  Tree regeneration following clearcut logging in red fir forests of California. , 1998 .

[31]  Patrick N. Halpin,et al.  Forest gradient response in Sierran landscapes: the physical template , 2000, Landscape Ecology.

[32]  Barbara J. Bond,et al.  Precision and accuracy of three alternative instruments for measuring soil water content in two forest soils of the Pacific Northwest , 2005 .

[33]  M. Barbour,et al.  SEEDLING GROWTH AND SURVIVAL OF RED AND WHITE FIR IN A SIERRA NEVADA ECOTONE , 1990 .

[34]  D. Binkley,et al.  Ecology and Management of Forest Soils , 2000 .

[35]  T. Bruns,et al.  Detection of plot-level changes in ectomycorrhizal communities across years in an old-growth mixed-conifer forest. , 2005, The New phytologist.

[36]  M. Allen,et al.  Ectomycorrhizae in a soil-weathered granitic bedrock regolith: linking matrix resources to plants , 2005 .

[37]  T. Stohlgren,et al.  Lodgepole pine (Pinus contorta) ecotones in Rocky Mountain National Park, Colorado, USA , 1997 .

[38]  R. B. Jackson,et al.  A global analysis of root distributions for terrestrial biomes , 1996, Oecologia.

[39]  Diana H. Wall,et al.  Interactions Underground: Soil biodiversity, mutualism, and ecosystem processes , 1999 .