Simulation of within-tree and between-tree shading of direct radiation in a forest canopy: effect of crown shape and sun elevation

Abstract The pattern of momentary within-tree and between-tree shading of foliage in direct radiation flux was simulated in fully-stocked model stands with a Poisson distribution of trees. The canopies were composed of umbrella-like (crown-height/crown-radius, or H/R = 3), average (H/R = 8) or columnar (H/R = 20) tree crowns. The momentary radiation-extinction probability was calculated for a large number of points (106−107 points per ha) within the canopy. The radiation-attenuation rate depended on the projected leaf-area density within the crown. At solar elevations of 5° and 45°, columnar crowns had the most between-tree and the least within-tree shading, while at a high sun elevation (85°) the same was true for umbrella-like crowns. At low and medium sun elevations (5° and 45°) the crown shape had a rather small effect on the total shading of foliage. At a high sun elevation, flat, horizontally extended crowns were clearly best-illuminated, because both within-tree and between-tree shadings were low.

[1]  H. Smolander,et al.  Photosynthesis of a Scots pine shoot: The effect of shoot inclination on the photosynthetic response of a shoot subjected to direct radiation , 1983 .

[2]  M. Cannell,et al.  Radiation Interception and Productivity of Willow , 1987 .

[3]  J. Monteith Climate and the efficiency of crop production in Britain , 1977 .

[4]  J. Monteith 2 – DOES LIGHT LIMIT CROP PRODUCTION? , 1981 .

[5]  W. G. Duncan Leaf Angles, Leaf Area, and Canopy Photosynthesis 1 , 1971 .

[6]  H. S. Horn The adaptive geometry of trees , 1971 .

[7]  Tõnu Oja,et al.  Regime of diffuse solar radiation in an individual norway spruce (Picea abies (L.) Karst.) Crown , 1984 .

[8]  T Kuuluvainen Crown architecture and stemwood production in Norway spruce (Picea abies (L.) Karst.). , 1988, Tree physiology.

[9]  S. M. Virmani,et al.  Crop productivity in relation to interception of photosynthetically active radiation , 1984 .

[10]  L. S. Jahnke,et al.  Influence of Photosynthetic Crown Structure on Potential Productivity of Vegetation, Based Primarily on Mathematical Models , 1965 .

[11]  T. Pukkala,et al.  Effect of crown shape and tree distribution on the spatial distribution of shade , 1987 .

[12]  Seppo Kellomäki,et al.  Theoretical computations on the role of crown shape in the absorption of light by forest trees , 1982 .

[13]  G. Mohren,et al.  Simulation of competition for light in even-aged stands of Douglas Fir , 1987 .

[14]  J. Ross The radiation regime and architecture of plant stands , 1981, Tasks for vegetation sciences 3.

[15]  T. Pukkala,et al.  Relationship between radiation interception and photosynthesis in forest canopies: effect of stand structure and latitude , 1989 .

[16]  Seppo Kellomäki,et al.  Effect of grouping of foliage on the within-stand and within-crown light regime: Comparison of random and grouping canopy models , 1983 .