How many Upper Triassic bone-bearing levels are there in Upper Silesia (southern Poland)? A critical overview of stratigraphy and facies

At least three widely separated bone-bearing intervals in the Upper Triassic succession of Upper Silesia, ranging in age from the Carnian to Rhaetian (i.e., in the interval of 25 Ma), are presented in papers by the Warsaw research group, led mainly by Jerzy Dzik and/or Grzegorz Niedźwiedzki. The stratigraphic arguments are reviewed for the vertebrate localities studied so far, in particular for the well-known middle Keuper sites at Krasiejow and Lipie Śląskie, to show that the previously proposed age assignments are still inadequately documented and questionable. This unreliability is exemplified by the evolving stratigraphic correlation of the fragmentary Silesian sections (8–18 m thick) with informal subsurface units from central-western Poland and with the German standard succession, ultimately not corroborated by comparison with the composite reference succession of the Upper Silesian Keuper, including new profiles (ca. 260 m thick) from the Woźniki K1 and Patoka 1 wells. Based on a multidisciplinary stratigraphic study covering consistent litho-, bio-, climato- and chemostratigraphic premises, focused on the regional reference section, two bone-bed levels only are recognized in the Patoka Marly Mudstone-Sandstone Member (= Steinmergelkeuper) of the Grabowa Formation, not very different in age (Classopollis meyeriana Palynozone; probably IVb Subzone): (1) the localized Krasiejow bone breccia level (early Norian in age) in the Opole region, and (2) the far more widely distributed Lisowice bone-bearing level (middle Norian) in a vast alluvial plain (braided to anastomosing river system) during the Eo-Cimmerian tectonic-pluvial episode. As a consequence of the principal uncertainties and controversies in Upper Triassic terrestrial stratigraphy, this is still a somewhat preliminary inference. Typical skeletal concentra- tions of a combined hydraulic/sedimentologic type, related to fluvial processes, are common in the Upper Silesian Fossil-Lagerstatten, although factors governing preservation are probably important, as well.

[1]  M. Antczak.  Late Triassic aetosaur (Archosauria) from Krasiejów (SW Poland): new species or an example of individual variation? , 2016 .

[2]  J. Zonneveld,et al.  Paleoenvironmental changes across the Carnian/Norian boundary in the Black Bear Ridge section, British Columbia, Canada , 2016 .

[3]  N. Konieczna,et al.  Nd and Sr isotopic evidence for provenance of clastic material of the Upper Triassic rocks of Silesia, Poland , 2015 .

[4]  D. Uhl,et al.  News from an old wood — Agathoxylon keuperianum (Unger) nov. comb. in the Keuper of Poland and France , 2015 .

[5]  G. Bachmann,et al.  The Germanic Triassic: correlations with the international chronostratigraphic scale, numerical ages and Milankovitch cyclicity , 2015 .

[6]  C. Pott,et al.  Coprolites of Late Triassic carnivorous vertebrates from Poland: an integrative approach , 2015 .

[7]  P. Skrzycki.  New Species of Lungfish (Sarcopterygii, Dipnoi) from the Late Triassic Krasiejów Site in Poland, with Remarks on the Ontogeny of Triassic Dipnoan Tooth Plates , 2015 .

[8]  M. Katz,et al.  Magnetostratigraphy, biostratigraphy, and chemostratigraphy of the Pignola-Abriola section: New constraints for the Norian-Rhaetian boundary , 2015 .

[9]  A. Mader.  Record of climatic changes in the Triassic palynological spectra from Poland , 2015 .

[10]  R. Butler,et al.  A New Species of Metoposaurus from the Late Triassic of Portugal and Comments on the Systematics and Biogeography of Metoposaurid Temnospondyls , 2015 .

[11]  K. Jewuła,et al.  Key aspects of the stratigraphy of the Upper Silesian middle Keuper, southern Poland , 2015 .

[12]  Idealized Model of Mineral Infillings in Bones of Fossil Freshwater Animals, on the Example of Late Triassic Metoposaurs from Krasiejów (Poland) , 2015 .

[13]  A. Becker.  Ambiguities in conchostracan biostratigraphy: A case study of the Permian–Triassic boundary , 2015 .

[14]  D. Uhl,et al.  EVIDENCE OF WILDFIRES DURING DEPOSITION OF THE UPPER SILESIAN KEUPER SUCCESSION, SOUTHERN POLAND , 2015 .

[15]  A. Mindszenty,et al.  Taphonomic and paleoecologic investigations of the Late Cretaceous (Santonian) Iharkút vertebrate assemblage (Bakony Mts, Northwestern Hungary) , 2015 .

[16]  J. Nawrocki,et al.  MAGNETIC POLARITY OF UPPER TRIASSIC SEDIMENTS OF THE GERMANIC BASIN IN POLAND , 2015 .

[17]  Anna Fijałkowska-Mader,et al.  Palynostratigraphy and palynofacies of the Upper Silesian Keuper (southern Poland) , 2015 .

[18]  S. Lucas.  Age and correlation of Late Triassic tetrapods from southern Poland , 2015 .

[19]  A. Fiorillo,et al.  DINOSAUR-BEARING HYPERCONCENTRATED FLOWS OF CRETACEOUS ARCTIC ALASKA: RECURRING CATASTROPHIC EVENT BEDS ON A DISTAL PALEOPOLAR COASTAL PLAIN , 2014 .

[20]  R. Butler,et al.  Basal dinosauriform and theropod dinosaurs from the mid–late Norian (Late Triassic) of Poland: implications for Triassic dinosaur evolution and distribution , 2014 .

[21]  G. Niedźwiedzki,et al.  Putative dicynodont coprolites from the Upper Triassic of Poland , 2014 .

[22]  J. Dzik,et al.  Skeletal Variation and Ontogeny of the Late Triassic Dinosauriform Silesaurus opolensis , 2014 .

[23]  G. Niedźwiedzki,et al.  Climatic reversals related to the Central Atlantic magmatic province caused the end-Triassic biotic crisis—Evidence from continental strata in Poland , 2014 .

[24]  L. Marynowski,et al.  Weathering, sedimentary and diagenetic controls of mineral and geochemical characteristics of the vertebrate-bearing Silesian Keuper , 2014, Clay Minerals.

[25]  N. Campione,et al.  Taphonomy of the Danek Bonebed: a monodominant Edmontosaurus (Hadrosauridae) bonebed from the Horseshoe Canyon Formation, Alberta , 2014 .

[26]  L. Krystyn,et al.  Towards accurate numerical calibration of the Late Triassic: High- precision U-Pb geochronology constraints on the duration of the Rhaetian , 2014 .

[27]  Grzegorz Pacyna.  Plant remains from the Polish Triassic. Present knowledge and future prospects , 2014 .

[28]  A. Feldman-Olszewska,et al.  A palaeoenvironmental reconstruction based on palynological analyses of Upper Triassic and Lower Jurassic sediments from the Holy Cross Mountains region , 2014 .

[29]  J. Nawrocki,et al.  Magnetostratigraphy of the Buntsandstein (Lower Triassic) in the Gorzów Wielkopolski IG 1 borehole, eastern German Basin in Poland: evidence of substantial diachronism of palynostratigraphic macrospore zones , 2014 .

[30]  M. Benton,et al.  Models for the Rise of the Dinosaurs , 2014, Current Biology.

[31]  R. Butler,et al.  Redescription of the phytosaurs Paleorhinus (‘Francosuchus’) angustifrons and Ebrachosuchus neukami from Germany, with implications for Late Triassic biochronology , 2014 .

[32]  V. Prokhorovich,et al.  PeRmIAn And TRIASSIc PAleonTology And STRATIgRAPhy of noRTh euRASIA , 2014 .

[33]  Stratigraphic Position of the Triassic Sediments in the Upper Silesian Region , 2014 .

[34]  T. Sulej,et al.  Mammal-Like Tooth from the Upper Triassic of Poland , 2013 .

[35]  T. Sulej,et al.  Aetosauria: a clade of armoured pseudosuchians from the Upper Triassic continental beds , 2013 .

[36]  A. Skawina.  Population dynamics and taphonomy of the Late Triassic (Carnian) freshwater bivalves from Krasiejów (Poland) , 2013 .

[37]  T. Marcinkiewicz,et al.  Miospores, megaspores and Lepidopteris ottonis (Goeppert) Schimper in the uppermost Triassic deposits from Poland , 2013 .

[38]  Nicole Klein,et al.  Unique growth pattern of Metoposaurus diagnosticus krasiejowensis (Amphibia, Temnospondyli) from the Upper Triassic of Krasiejów, Poland , 2013 .

[39]  S. Lucas.  PLANT MEGAFOSSIL BIOSTRATIGRAPHY AND BIOCHRONOLOGY, UPPER TRIASSIC CHINLE GROUP, WESTERN USA , 2013 .

[40]  Grzegorz Sadlok,et al.  Upper Triassic vertebrate tracks from Kraków–Częstochowa Upland, southern Poland , 2013 .

[41]  M. Menning,et al.  Erläuterung zur Stratigraphischen Tabelle von Deutschland Kompakt 2012 [Explanatory notes to the Stratigraphic Table of Germany Compact 2012] , 2012 .

[42]  D. Konietzko-Meier,et al.  Histological characteristics of the vertebral intercentra of Metoposaurus diagnosticus (Temnospondyli) from the Upper Triassic of Krasiejów (Upper Silesia, Poland) , 2012, Earth and Environmental Science Transactions of the Royal Society of Edinburgh.

[43]  T. Sulej,et al.  A New Late Triassic Vertebrate Fauna from Poland with Turtles, Aetosaurs, and Coelophysoid Dinosaurs , 2012 .

[44]  A. Bodzioch,et al.  Unraveling the origin of the Late Triassic multitaxic bone accumulation at Krasiejów (S Poland) by diagenetic analysis , 2012 .

[45]  L. Tanner,et al.  The Late Triassic timescale: Age and correlation of the Carnian–Norian boundary , 2012 .

[46]  D. Mukherjee,et al.  Taphonomy of an Upper Triassic vertebrate bonebed: A new rhynchosaur (Reptilia; Archosauromorpha) accumulation from India , 2012 .

[47]  J. Dzik,et al.  A Large Predatory Archosaur from the Late Triassic of Poland , 2012 .

[48]  J. Dzik,et al.  Umbonal musculature and relationships of the Late Triassic filibranch unionoid bivalves , 2011 .

[49]  H. Kozur,et al.  A latest Norian age for insect-bearing beds of the Fuchsberg and Langenberg near Seinstedt, northern foreland of the Harz Mountains Lower Saxony, Germany , 2011 .

[50]  P. Gorzelak,et al.  Bite traces on dicynodont bones and the early evolution of large terrestrial predators , 2011 .

[51]  S. Lucas,et al.  Taphonomy of the Lamy Amphibian Quarry: A Late Triassic bonebed in New Mexico, U.S.A. , 2010 .

[52]  T. Sulej,et al.  A new dicynodont–archosaur assemblage from the Late Triassic (Carnian) of Poland , 2010, Earth and Environmental Science Transactions of the Royal Society of Edinburgh.

[53]  R. Irmis.  Evaluating hypotheses for the early diversification of dinosaurs , 2010, Earth and Environmental Science Transactions of the Royal Society of Edinburgh.

[54]  J. Dzik,et al.  The Axial Skeleton of Silesaurus opolensis , 2010 .

[55]  M. Norell,et al.  The origin and early radiation of dinosaurs , 2010 .

[56]  H. Sues,et al.  Triassic Life on Land: The Great Transition , 2010 .

[57]  Andrea Anelli,et al.  Environmental control on the end of the Dolomia Principale/Hauptdolomit depositional system in the central Alps: Coupling sea-level and climate changes , 2010 .

[58]  P. Wignall,et al.  Triassic climates — State of the art and perspectives , 2010 .

[59]  T. Sulej.  The skull of an early Late Triassic aetosaur and the evolution of the stagonolepidid archosaurian reptiles , 2010 .

[60]  B. Gruszka,et al.  Evidence for a very low-energy fluvial system: a case study from the dinosaur-bearing Upper Triassic rocks of Southern Poland , 2010 .

[61]  K. Angielczyk,et al.  Ecologically distinct dinosaurian sister group shows early diversification of Ornithodira , 2010, Nature.

[62]  H. Kozur,et al.  The biostratigraphic importance of conchostracans in the continental Triassic of the northern hemisphere , 2010 .

[63]  G. Racki.  Ewolucja środowisk lądowych kajpru Górnego Śląska jako biotopów kręgowców-nowy projekt badawczy , 2010 .

[64]  M. A. Salamon,et al.  Kopalne Biocenozy w Czasie i Przestrzeni , 2010 .

[65]  S. Cirilli.  Upper Triassic–lowermost Jurassic palynology and palynostratigraphy: a review , 2010 .

[66]  S. Lucas.  The Triassic timescale based on nonmarine tetrapod biostratigraphy and biochronology , 2010 .

[67]  Wolfram M Kürschner,et al.  Triassic palynology of central and northwestern Europe: a review of palynofloral diversity patterns and biostratigraphic subdivisions , 2010 .

[68]  G. Sadlok,et al.  BITE MARKS ON AN UPPER TRIASSIC DICYNODONTID TIBIA FROM ZAWIERCIE, KRAKÓW-CZĘSTOCHOWA UPLAND, SOUTHERN POLAND , 2010 .

[69]  B. Simoneit,et al.  WIDESPREAD UPPER TRIASSIC TO LOWER JURASSIC WILDFIRE RECORDS FROM POLAND: EVIDENCE FROM CHARCOAL AND PYROLYTIC POLYCYCLIC AROMATIC HYDROCARBONS , 2009 .

[70]  Rodney D. Scheetz,et al.  Taphonomy of debris-flow hosted dinosaur bonebeds at Dalton Wells, Utah (Lower Cretaceous, Cedar Mountain Formation, USA) , 2009 .

[71]  P. Gorzelak,et al.  Pathologies of non-marine bivalve shells from the Late Triassic of Poland , 2009 .

[72]  Alan G. Stevenson,et al.  Petroleum geological atlas of the Southern Permian Basin area , 2009 .

[73]  R. Butler,et al.  The Taxonomy and Anatomy of Rauisuchian Archosaurs from the Late Triassic of Germany and Poland , 2009 .

[74]  A. Seilacher.  Biomats, biofilms, and bioglue as preservational agents for arthropod trackways , 2008 .

[75]  J. Dzik,et al.  A Dicynodont-Theropod Association in the Latest Triassic of Poland , 2008 .

[76]  J. Dzik.  Gill structure and relationships of the Triassic cycloid crustaceans , 2008, Journal of morphology.

[77]  T. McCann,et al.  The Geology of Central Europe - Volume 1 Precambrian and Palaeozoic , 2008 .

[78]  N. Tougiannidis,et al.  Orbital control on Upper Triassic Playa cycles of the Steinmergel-Keuper (Norian): A new concept for ancient playa cycles , 2008 .

[79]  M. Franz.  Litho- und Leitflächenstratigraphie, Chronostratigraphie, Zyklo- und Sequenzstratigraphie des Keupers im östlichen zentraleuropäischen Becken (Deutschland, Polen) und dänischen Becken (Dänemark, Schweden) , 2008 .

[80]  Ewa Barycka.  Morphology and ontogeny of the humerus of the Triassic temnospondyl amphibian Metoposaurus diagnosticus , 2007 .

[81]  J. Dzik,et al.  A REVIEW OF THE EARLY LATE TRIASSIC KRASIEJÓW BIOTA FROM SILESIA , POLAND , 2007 .

[82]  A. Behrensmeyer.  Bonebeds through Time , 2007 .

[83]  J. Szulc.  Stop III. 1. Laryszów : communal waste depot , 2007 .

[84]  S. Kidwell,et al.  A Conceptual Framework for the Genesis and Analysis of Vertebrate Skeletal Concentrations , 2007 .

[85]  S. Lucas,et al.  BIOCHRONOLOGICAL SIGNIFICANCE OF LATE TRIASSIC TETRAPODS FROM KRASIEJÓW , POLAND , 2007 .

[86]  J. Szulc,et al.  The Upper Triassic crenogenic limestones in Upper Silesia (southern Poland) and their paleoenvironmental context , 2006 .

[87]  Edgar Nitsch,et al.  Sedimente und Sedimentationspausen im deutschen Keuper: Wie vollständig ist die Überlieferung der späten Triaszeit? , 2005 .

[88]  E. Sienkiewicz,et al.  Late Triassic charophytes around the bone-bearing bed at Krasiejów (SW Poland) - palaeoecological and environmental remarks , 2005 .

[89]  G. Arp,et al.  Palaeoenvironmental significance of lacustrine stromatolites of the Arnstadt Formation (“Steinmergelkeuper”, Upper Triassic, N-Germany) , 2005 .

[90]  R. Rogers.  Fine-grained debris flows and extraordinary vertebrate burials in the Late Cretaceous of Madagascar , 2005 .

[91]  Edgar Nitsch,et al.  Paläoflora und Ablagerungsräume im unterfränkischen Keuper (Exkursion H am 1. April 2005) , 2005 .

[92]  T. Sulej.  A NEW RAUISUCHIAN REPTILE (DIAPSIDA: ARCHOSAURIA) FROM THE LATE TRIASSIC OF POLAND , 2005 .

[93]  J. Szulc.  Sedimentary environments of the vertebrate-bearing Norian deposits from Krasiejow, Upper Silesia (Poland) Vertebraten-führende sedimentäre Ablagerungsbereiche im Norium (Obere Trias) von Krasiejow, Oberschlesien (Polen) , 2005 .

[94]  E. Olempska.  Late Triassic spinicaudatan crustaceans from Southwestern Poland , 2004 .

[95]  J. Dzik.  A BEAKED HERBIVOROUS ARCHOSAUR WITH DINOSAUR AFFINITIES FROM THE EARLY LATE TRIASSIC OF POLAND , 2003 .

[96]  Andrew S. Cohen,et al.  Paleolimnology: The History and Evolution of Lake Systems , 2004 .

[97]  T. Sulej.  Species discrimination of the Late Triassic temnospondyl amphibian Metoposaurus diagnosticus , 2002 .

[98]  M. Ryan,et al.  The Taphonomy of a Centrosaurus (Ornithischia: Certopsidae) Bone Bed from the Dinosaur Park Formation (Upper Campanian), Alberta, Canada, with Comments on Cranial Ontogeny , 2001 .

[99]  J. Dzik.  A New Paleorhinus Fauna in the Early Late Triassic of Poland , 2001 .

[100]  B. Makaske.  Anastomosing rivers: a review of their classification, origin and sedimentary products , 2001 .

[101]  W. Ricken,et al.  The stratigraphic and geochemical record of Playa Cycles: monitoring a Pangaean monsoon-like system (Triassic, Middle Keuper, S. Germany) , 2000 .

[102]  François Therrien,et al.  Paleoenvironments of Early Theropods, Chinle Formation (Late Triassic), Petrified Forest National Park, Arizona , 2000 .

[103]  S. Kidwell,et al.  Associations of Vertebrate Skeletal Concentrations and Discontinuity Surfaces in Terrestrial and Shallow Marine Records: A Test in the Cretaceous of Montana , 2000, The Journal of Geology.

[104]  J. Szulc.  MIDDLE TRIASSIC EVOLUTION OF THE NORTHERN PERI-TETHYS AREA AS INFLUENCED BY EARLY OPENING OF THE TETHYS OCEAN , 2000 .

[105]  Bohacs,et al.  Abstract: Lake-Basin Type, Source Potential, and Hydrocarbon Character: An Integrated Sequence-Stratigraphic-Geochemical Framework , 1999 .

[106]  S. Lucas.  Global Triassic tetrapod biostratigraphy and biochronology , 1998 .

[107]  F. Jenkins,et al.  A Late Triassic lake system in East Greenland: facies, depositional cycles and palaeoclimate , 1998 .

[108]  A. Behrensmeyer,et al.  Taphonomy and Time Resolution of Bone Assemblages in a Contemporary Fluvial System: The East Fork River, Wyoming , 1996 .

[109]  J. V. Couvering,et al.  Global Geological Record of Lake Basins, Volume 1 , 1996 .

[110]  S. Squillace,et al.  Environmental control , 1992, Otolaryngology--head and neck surgery : official journal of American Academy of Otolaryngology-Head and Neck Surgery.

[111]  Richard G. Thomas,et al.  Fluvial processes and vertebrate taphonomy: the upper cretaceous Judith River formation, South-Central dinosaur Provincial Park, Alberta, Canada , 1988 .

[112]  Soumen Sarkar.  Petrology of caliche-derived peloidal calcirudite/ calcarenite in the late triassic maleri formation of the pranhita-Godavari valley, South India , 1988 .

[113]  A. Behrensmeyer.  Vertebrate preservation in fluvial channels , 1988 .

[114]  R. V. Demicco,et al.  Facies sequences of a semi-arid closed basin: the Lower Jurassic East Berlin Formation of the Hartford Basin, New England, U.S.A. , 1986 .

[115]  A. Seilacher,et al.  Sedimentological, ecological and temporal patterns of fossil Lagerstätten , 1985 .

[116]  T. Orłowska-Zwolińska.  Palynological zones of the Polish epicontinental triassic , 1985 .

[117]  M. Kraus,et al.  Vertebrate fossil-bearing paleosol units (Willwood Formation, Lower Eocene, Northwest Wyoming, U.S.A.): Implications for taphonomy, biostratigraphy, and assemblage analysis , 1981 .

[118]  Jiannan Zhao,et al.  Lakes , 2021, Springer New York.

[119]  Prussia,et al.  Geologie von Oberschlesien. , 1870 .