Lower Miocene Cervoid Ruminants (Artiodactyla) from the Koura Formation, Matsue, Southwestern Japan

Abstract. Dental fossils of an indeterminate small cervoid (Ruminantia, Artiodactyla, Mammalia) from the lower Miocene Koura Formation of southwestern Japan are described. The examined specimens of Cervoidea fam., gen. et sp. indet. consist of selenodont lower molars with plesiomorphic characteristics of early cervoids: a strong external postprotocristid (or Palaeomeryx-fold), a deep groove on the lingual wall of the internal postmetacristid, and opening posterior cristids between the postentocristid and the posthypocristid. The lower molars are similar in shape and size with those of Miocene Cervidae rather than Moschidae or Palaeomerycidae because the present specimens are comparable to those of Dicrocerus tokunagai, which is one of the lower/ middle Miocene cervids from northeastern Japan, and D. grangeri from the middle Miocene of China. The lower Miocene Nakamura Formation (ca. 19 Ma) of central Japan also yielded a small cervoid species sharing the lower molar characteristics with Cervoidea fam., gen. et sp. indet. from the Koura Formation, suggesting a faunal or biostratigraphical correlation between the two formations.

[1]  Tingjiang Peng,et al.  Diversity of Moschidae (Ruminantia, Artiodactyla, Mammalia) in the Middle Miocene of China , 2015 .

[2]  K. Miyata,et al.  Miocene Land Mammals and Stratigraphy of Japan , 2013 .

[3]  Gertrud E. Rössner,et al.  Dental nomenclature in Ruminantia: Towards a standard terminological framework , 2011 .

[4]  S. Kawano,et al.  X-Ray Computed Tomography Examination of a Fossil Beaver Tooth from the Lower Miocene Koura Formation of Western Japan , 2011 .

[5]  J. Morales,et al.  The genus Hispanomeryx (Mammalia, Ruminantia, Moschidae) and its bearing on musk deer phylogeny and systematics , 2010 .

[6]  E. Douzery,et al.  Molecular and morphological phylogenies of ruminantia and the alternative position of the moschidae. , 2003, Systematic biology.

[7]  T. Shikama,et al.  Note on an Occurrence of Dicrocerus in-the Daigo Group of the Ibaraki Prefecture, Japan , 1952 .

[8]  Y. Ando,et al.  A cervoid tooth from the lower Miocene Nakamura Formation of the Mizunami Group in Kani City, Gifu Prefecture, central Japan , 2016 .

[9]  A. Furukawa,et al.  Stepwise environmental changes in the lower Miocene Koura Formation, southwest Japan, associated with Japan Sea evolution , 2013 .

[10]  I. Vislobokova Eocene - early Miocene ruminants in Asia , 1997 .

[11]  C. Janis,et al.  The interrelationships of higher ruminant families : with special emphasis on the members of the Cervoidea. American Museum novitates ; no. 2893 , 1987 .

[12]  J. Sudre,et al.  Relations phylétiques de Bachitherium Filhol, ruminant de l'Oligocène d'Europe occidentale , 1987 .

[13]  Hikoshichiro . Matsumoto On a New Archetypal Fossil Cervid from the Prov. of Mino , 1918 .